Khodzhakul Formation

The Khodzhakul Formation is a Cenomanian aged geologic formation in Uzbekistan. Dinosaur remains are among the fossils that have been recovered from the formation.[1] As well as those of Mammals.[2] It is part of the same stratigraphic succession as the overlying Bissekty Formation.

Khodzhakul Formation Stratigraphic range: Cenomanian ~93–89 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Type	Geological formation
Underlies	Tamdytruba & Bissekty Formations
Overlies	Bortesken Formation
Lithology
Primary	Sandstone
Location
Coordinates	40.1°N 65.8°E
Approximate paleocoordinates	35.9°S 65.5°E
Region	Karakalpakstan
Country	Uzbekistan
Khodzhakul Formation (Uzbekistan)

Fossil content

One of the largest mammals of the Mesozoic, Oxlestes, occurred in this formation, and may have predated on dinosaur species.[3]

Dinosaurs
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Asiaceratops[4]	A. salsopaludalis[4]		Upper[4]	"Teeth, cranial fragments, phalanx."[5]
Gilmoreosaurus[6]	G. atavus[6]		Upper[4]
Horezmavis[7]	H. eocretaceous[7]		Lower[7]	"Tarsometatarsus."[8]
Kulceratops[4]	K. kulensis[4]		Upper[4]	"Fragmentary material."[5]
Therizinosauroidea	Indeterminate[9]		Upper
Pectinodon[4]	P. Sp[4]		Upper[4]

Mammals[2]
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Bobolestes	B. zenge			Maxilla and dentary fragments	Zalambdalestoidea
Eozhelestes	E. mangit			dentary fragments and teeth
Sheikhdzheilia	S. rezvyii			Maxilla and dentary fragments	?Zhelestidae
Zalambdalestidae	Indeterminate			Right petrosal
‘‘Zhelestidae’’	Indeterminate			Left dentary and dentary fragment	Large taxon

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon		Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

See also

• List of dinosaur-bearing rock formations

References

1. Weishampel, et al., 2004, "Dinosaur distribution." pp.517-607
2. Averianov, Alexander; Archibald, J. David (August 2005). "Mammals from the mid-Cretaceous Khodzhakul Formation, Kyzylkum Desert, Uzbekistan". Cretaceous Research. 26 (4): 593–608. doi:10.1016/j.cretres.2005.03.007.
3. Michael J. Benton,Mikhail A. Shishkin, David M. Unwin, The Age of Dinosaurs in Russia and Mongolia, Cambridge University Press, 04/12/2003 - 740 páginas
4. "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Upper Khodzhakul Formation," in Weishampel, et al. (2004). Page 594.
5. "Table 22.1," in Weishampel, et al. (2004). Page 480.
6. Listed as "? Gilmoreosaurus sp." in "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Upper Khodzhakul Formation," in Weishampel, et al. (2004). Page 594.
7. "40.3 Qoraqalpoghiston Respublikasi, Uzbekistan; 1. Lower Khodzhakul Formation," in Weishampel, et al. (2004). Page 563.
8. "Table 11.1," in Weishampel, et al. (2004). Page 214.
9. Sues, Hans-Dieter; Averianov, Alexander (2016). "Therizinosauroidea (Dinosauria: Theropoda) from the Upper Cretaceous of Uzbekistan". Cretaceous Research. 59: 155–178. doi:10.1016/j.cretres.2015.11.003.

Bibliography

• Weishampel, David B.; Peter Dodson, and Halszka Osmólska (eds.). 2004. The Dinosauria, 2nd edition, 1–880. Berkeley: University of California Press. Accessed 2019-02-21. ISBN 0-520-24209-2