List of Cyphophthalmi species

This page lists all described species of the harvestman suborder Cyphophthalmi. Unless otherwise noted, information is taken from Giribet's 2000 taxonomic catalogue[1] and from Giribet et al., 2012.[2]

Main article: Cyphophthalmi

Boreophthalmi

Sironidae
  • Cyphophthalmus Joseph, 1868[3]
  • Cyphophthalmus beschkovi (Mitov, 1994)
  • Cyphophthalmus bithynicus (Gruber, 1969)
  • Cyphophthalmus conocephalus Karaman, 2009
  • Cyphophthalmus corfuanus (Kratochvíl, 1938)
  • Cyphophthalmus duricorius Joseph, 1868
  • Cyphophthalmus eratoae (Juberthie, 1968)
  • Cyphophthalmus ere Karaman, 2008
  • Cyphophthalmus gjorgjevici (Hadži, 1933)
  • Cyphophthalmus gordani Karaman, 2009
  • Cyphophthalmus hlavaci Karaman, 2009
  • Cyphophthalmus klisurae (Hadži, 1973)
  • Cyphophthalmus kratochvili Karaman, 2009
  • Cyphophthalmus markoi Karaman, 2008
  • Cyphophthalmus martensi Karaman, 2009
  • Cyphophthalmus minutus (Kratochvíl, 1938)
  • Cyphophthalmus neretvanus Karaman, 2009
  • Cyphophthalmus noctiphilus (Kratochvíl, 1940)
  • Cyphophthalmus nonveilleri Karaman, 2008
  • Cyphophthalmus ognjenovici Karaman, 2009
  • Cyphophthalmus ohridanus (Hadži, 1973)
  • Cyphophthalmus paradoxus (Kratochvíl, 1958)
  • Cyphophthalmus paragamiani Karaman, 2009
  • Cyphophthalmus rumijae Karaman, 2009
  • Cyphophthalmus serbicus (Hadži, 1973)
  • Cyphophthalmus silhavyi (Kratochvíl, 1938)
  • Cyphophthalmus solentiensis Dreszer, Raða & Giribet, 2015[4]
  • Cyphophthalmus teyrovskyi (Kratochvíl, 1938)
  • Cyphophthalmus thracicus Karaman, 2009
  • Cyphophthalmus trebinjanus Karaman, 2009
  • Cyphophthalmus yalovensis (Gruber, 1969)
  • Cyphophthalmus zetae Karaman, 2009
  • Iberosiro de Bivort & Giribet, 2004[5]
  • Iberosiro distylos de Bivort & Giribet, 2004
  • Iberosiro rosae Giribet, Merino-Sáinz & Benavides, 2017
  • Marwe Shear, 1985
  • Marwe coarctata Shear, 1985
  • Odontosiro Juberthie, 1961
  • Odontosiro lusitanicus Juberthie, 1961
  • Paramiopsalis Juberthie, 1962[5]
  • Paramiopsalis anadonae Giribet, Merino-Sáinz & Benavides, 2017
  • Paramiopsalis eduardoi Murienne & Giribet, 2009
  • Paramiopsalis ramblae Benavides & Giribet, 2017
  • Paramiopsalis ramulosus Juberthie, 1962
  • Parasiro Hansen and Sørensen, 1904
  • Parasiro coiffaiti Juberthie, 1956
  • Parasiro corsicus (Simon, 1872)
  • Parasiro minor Juberthie, 1958
  • Suzukielus Juberthie, 1970
  • Suzukielus sauteri (Roewer, 1916)

Fangensinae
  • Fangensis Rambla, 1994
  • Fangensis cavernarus Schwendinger & Giribet, 2005
  • Fangensis leclerci Rambla, 1994
  • Fangensis spelaeus Schwendinger & Giribet, 2005
  • Giribetia Clouse, 2012
  • Giribetia insulanus (Schwendinger & Giribet, 2005)

Leptopsalidinae[7]
  • Leptopsalis Thorell, 1882
  • Leptopsalis beccarii Thorell, 1882
  • Leptopsalis dumoga (Shear, 1993)
  • Leptopsalis foveolata Clouse & Schwendinger, 2012[8]
  • Leptopsalis hillyardi (Shear, 1993)
  • Leptopsalis javana (Thorell, 1882)
  • Leptopsalis laevichelis (Roewer, 1942)
  • Leptopsalis lydekkeri (Clouse & Giribet, 2007)
  • Leptopsalis modesta (Hansen & Sørensen, 1904)
  • Leptopsalis novaguinea (Clouse & Giribet, 2007)
  • Leptopsalis pangrango (Shear, 1993)
  • Leptopsalis ramblae (Giribet, 2002)
  • Leptopsalis sedgwicki (Shear, 1979)
  • Leptopsalis sulcata (Hansen & Sørensen, 1904)
  • Leptopsalis tambusisi (Shear, 1993)
  • Leptopsalis thorellii (Hansen & Sørensen, 1904)
  • Leptopsalis weberii (Hansen & Sørensen, 1904)
  • Miopsalis Thorell, 1890
  • Miopsalis collinsi (Shear, 1993)
  • Miopsalis dillyi Schmidt, Clouse & Sharma, 2020[9]
  • Miopsalis globosa (Schwendinger & Giribet, 2004)
  • Miopsalis gryllospeca (Shear, 1993)
  • Miopsalis kinabalu (Shear, 1993)
  • Miopsalis leakeyi (Shear, 1993)
  • Miopsalis lionota (Pocock, 1897)
  • Miopsalis mulu (Shear, 1993)
  • Miopsalis pocockii (Hansen & Sørensen, 1904)
  • Miopsalis pulicaria Thorell, 1890
  • Miopsalis sabah (Shear, 1993)
  • Miopsalis silhavyi (Rambla, 1991)
  • Miopsalis tarumpitao (Shear, 1993)

Stylocellinae
  • Meghalaya annandalei Giribet, Sharma, & Bastawade, 2007
  • Stylocellus Westwood, 1874
  • Stylocellus lornei Clouse, 2012
  • Stylocellus sumatranus Westwood, 1874

Scopulophthalmi

Pettalidae
  • Aoraki calcarobtusa (Forster, 1952)
  • Aoraki calcarobtusa calcarobtusa (Forster, 1952)
  • Aoraki calcarobtusa westlandica (Forster, 1952)
  • Aoraki crypta (Forster, 1948)
  • Aoraki denticulata (Forster, 1948)
  • Aoraki denticulata denticulata (Forster, 1948)
  • Aoraki denticulata major (Forster, 1948)
  • Aoraki granulosa (Forster, 1952)
  • Aoraki healyi (Forster, 1948)
  • Aoraki inerma (Forster, 1948)
  • Aoraki inerma inerma (Forster, 1948)
  • Aoraki inerma stephenesis (Forster, 1952)
  • Aoraki longitarsa (Forster, 1952)
  • Aoraki tumidata (Forster, 1948)
  • Austropurcellia Juberthie, 1988[11]
  • Austropurcellia absens Boyer & Popkin-Hall, 2015
  • Austropurcellia acuta Popkin-Hall & Boyer, 2014
  • Austropurcellia alata Boyer & Reuter, 2012
  • Austropurcellia arcticosa (Cantrell, 1980)
  • Austropurcellia barbata Popkin-Hall & Boyer, 2014
  • Austropurcellia cadens Baker & Boyer, 2015
  • Austropurcellia capricornia (Davies, 1977)
  • Austropurcellia clousei Boyer, Baker & Popkin-Hall, 2015
  • Austropurcellia culminis Boyer & Reuter, 2012
  • Austropurcellia daviesae (Juberthie, 1989)
  • Austropurcellis despectata Boyer & Reuter, 2012
  • Austropurcellia finniganensis Popkin-Hall, Jay & Boyer, 2016
  • Austropurcellia forsteri (Juberthie, 2000)
  • Austropurcellia fragosa Popkin-Hall, Jay & Boyer, 2016
  • Austropurcellia giribeti Boyer & Quay, 2015
  • Austropurcellia megatanka Jay, Coblens & Boyer, 2016
  • Austropurcellia monteithi Jay, Popkin-Hall, Coblens & Boyer, 2016
  • Austropurcellia riedeli Jay, Oberski & Boyer, 2016
  • Austropurcellia nuda Popkin-Hall, Jay & Boyer, 2016
  • Austropurcellia scoparia Juberthie, 1988
  • Austropurcellia sharmai Boyer & Quay, 2015
  • Austropurcellia superbensis Popkin-Hall & Boyer, 2014
  • Austropurcellia tholei Baker & Boyer, 2015
  • Austropurcellia vicina Boyer & Reuter, 2012
  • Austropurcellia woodwardi (Forster, 1955)
  • Chileogovea Roewer, 1961
  • Chileogovea jocasta Shear, 1993
  • Chileogovea oedipus Roewer, 1961
  • Karripurcellia Giribet, 2003[12]
  • Karripurcellia harveyi Giribet, 2003
  • Karripurcellia peckorum Giribet, 2003
  • Karripurcellia sierwaldae Giribet, 2003
  • Manangotria Shear & Gruber, 1996
  • Manangotria taolanaro Shear & Gruber, 1996
  • Neopurcellia Forster, 1948[10]
  • Neopurcellia salmoni Forster, 1948
  • Parapurcellia Rosas Costa, 1950[13]
  • Parapurcellia amatola de Bivort & Giribet, 2010
  • Parapurcellia convexa de Bivort & Giribet, 2010
  • Parapurcellia fissa (Lawrence, 1939)
  • Parapurcellia minuta de Bivort & Giribet, 2010
  • Parapurcellia monticola (Lawrence, 1939)
  • Parapurcellia natalia de Bivort & Giribet, 2010
  • Parapurcellia rumpiana (Lawrence, 1933)
  • Parapurcellia peregrinator (Lawrence, 1963)
  • Parapurcellia silvicola (Lawrence, 1939)
  • Parapurcellia staregai de Bivort & Giribet, 2010
  • Parapurcellia transvaalica (Lawrence, 1963)[14]
  • Pettalus Thorell, 1876
  • Pettalus brevicauda Pocock, 1897
  • Pettalus cimiciformis (Cambridge, 1875)
  • Pettalus lampetides Sharma & Giribet 2006[15]
  • Pettalus thwaitesi Sharma, Karunarathna & Giribet, 2009[16]
  • Purcellia Hansen & Sørensen, 1904[13]
  • Purcellia argasiformis (Lawrence, 1931)[14]
  • Purcellia griswaldi de Bivort & Giribet, 2010
  • Purcellia illustrans Hansen and Sørensen, 1904
  • Purcellia lawrencei de Bivort & Giribet, 2010
  • Purcellia leleupi Staręga, 2008[17]
  • Rakaia antipodiana Hirst, 1925
  • Rakaia collaria Roewer, 1942
  • Rakaia dorothea Phillips and Grimmett, 1932
  • Rakaia florensis (Forster, 1948)
  • Rakaia isolata Forster, 1952
  • Rakaia lindsayi Forster, 1952
  • Rakaia macra Boyer and Giribet, 2003
  • Rakaia magna Forster, 1948
  • Rakaia magna australis Forster, 1952
  • Rakaia magna magna Forster, 1948
  • Rakaia media Forster, 1948
  • Rakaia media media Forster, 1948
  • Rakaia media insula Forster, 1952
  • Rakaia minutissima (Forster, 1948)
  • Rakaia pauli Forster, 1952
  • Rakaia solitaria Forster, 1948
  • Rakaia sorenseni Forster, 1952
  • Rakaia sorenseni digitata Forster, 1952
  • Rakaia sorenseni sorenseni Forster, 1952
  • Rakaia stewartiensis Forster, 1948
  • Rakaia uniloca Forster, 1952

Sternophthalmi

Neogoveidae[18]
  • Brasiliogovea Martens, 1969
  • Brasiliogovea aphantostylus Benavides, Hormiga, & Giribet, 2019
  • Brasiliogovea chiribiqueta Benavides & Giribet, 2013
  • Brasiliogovea microphaga Martens, 1969
  • Brasiliogovea microstylus Benavides, Hormiga, & Giribet, 2019
  • Brasiliogovea yacambuensis Benavides, Hormiga, & Giribet, 2019
  • Canga DaSilva, Pinto-da-Rocha & Giribet, 2010
  • Huitaca Shear, 1979
  • Huitaca boyacanensis Benavides & Giribet, 2013
  • Huitaca bitaco Benavides & Giribet, 2013
  • Huitaca caldas Benavides & Giribet, 2013
  • Huitaca depressa Benavides & Giribet, 2013
  • Huitaca sharkeyi Benavides & Giribet, 2013
  • Huitaca tama Benavides & Giribet, 2013
  • Huitaca ventralis Shear, 1979
  • Leggogovia Benavides & Giribet, 2019
  • Leggogovia pabsgarnoni (Legg. 1990)
  • Metagovea Rosas-Costa, 1950
  • Metagovea disparunguis Rosas Costa, 1950
  • Metagovea ligiae Giupponi & Kury, 2015
  • Metagovea matapi Benavides, Hormiga, & Giribet, 2019
  • Metagovea philipi Goodnight & Goodnight, 1980
  • Metagovea planada Benavides, Hormiga, & Giribet, 2019
  • Metasiro americanus (Davis, 1933)
  • Metasiro savannahensis Clouse & Wheeler, 2014
  • Metasiro sassafrasensis Clouse & Wheeler, 2014
  • Microgovia Benavides, Hormiga, & Giribet, 2019
  • Microgovia chenepau Benavides, Hormiga, & Giribet, 2019
  • Microgovia oviformis Martens, 1969
  • Neogovea Hinton, 1938
  • Neogovea branstetteri Benavides, Hormiga, & Giribet, 2019
  • Neogovea enigmatica Martens, 2019
  • Neogovea hormigai Benavides & Giribet, 2013
  • Neogovea immsi Hinton, 1938
  • Neogovea kamakusa Shear, 1977
  • Neogovea kartabo Shear, 1977
  • Neogovea matawai Benavides, Hormiga, & Giribet, 2019
  • Neogovea virginie Jocqué & Jocqué, 2011
  • Parogovia Hansen, 1921
  • Parogovia gabonica (Juberthie, 1969)
  • Parogovia montealensis Benavides & Giribet, 2019
  • Parogovia parasironoides Hiřman, Kotyk, Kotyková Varadínová,& Šťáhlavský, 2018
  • Parogovia prietoi Benavides & Giribet, 2019
  • Parogovia putnami Benavides & Giribet, 2019
  • Parogovia sironoides Hansen, 1921
  • Tucanogovea Karaman, 2014
  • Tucanogovea schusteri Karaman, 2014
  • Waiwaigovia Benavides, Hormiga, & Giribet, 2019
  • Waiwaigovia shultzi Benavides, Hormiga, & Giribet, 2019

Ogoveidae[20]
  • Ogovea cameroonensis Giribet & Prieto, 2003
  • Ogovea grossa (Hansen & Sørensen, 1904)
  • Ogovea nasuta (Hansen, 1921)

Troglosironidae[21]
  • Troglosiro aelleni Juberthie, 1979
  • Troglosiro brevifossa Sharma & Giribet, 2009
  • Troglosiro juberthiei Shear, 1993
  • Troglosiro longifossa Sharma & Giribet, 2005
  • Troglosiro monteithi Sharma & Giribet, 2009
  • Troglosiro ninqua Shear, 1993
  • Troglosiro oscitatio Sharma & Giribet, 2009
  • Troglosiro platnicki Shear, 1993
  • Troglosiro raveni Shear, 1993
  • Troglosiro sheari Sharma & Giribet, 2009
  • Troglosiro tillierorum Shear, 1993
  • Troglosiro urbanus Sharma & Giribet, 2009
  • Troglosiro wilsoni Sharma & Giribet, 2009

Incertae sedis
  • Ankaratra franzi Shear & Gruber, 1996
  • Shearogovea Giribet, 2011
  • Shearogovea mexasca (Shear, 1977)

References
  1. Giribet, Gonzalo (2000). "Catalogue of the Cyphophthalmi of the World (Arachnida, Opiliones)" (PDF). Revista Ibérica de Aracnología. 2: 49–76.
  2. Giribet, Gonzalo; Sharma, Prashant; Benavides, Ligia; et al. (2012). "Evolutionary and biogeographical history of an ancient and global group of arachnids (Arachnida: Opiliones: Cyphophthalmi) with a new taxonomic arrangement". Biological Journal of the Linnean Society. 105 (1): 92–130. doi:10.1111/j.1095-8312.2011.01774.x.
  3. Karaman, Ivo M. (2009-06-01). "The taxonomical status and diversity of Balkan sironids (Opiliones, Cyphophthalmi) with descriptions of twelve new species". Zoological Journal of the Linnean Society. 156 (2): 260–318. doi:10.1111/j.1096-3642.2009.00446.x. ISSN 0024-4082.
  4. Giribet, Gonzalo; Raða, Tonći; Dreszer, Taras B. (2015). "Cyphophthalmus solentiensis sp. nov. (Cyphophthalmi, Sironidae), a New Endogean Mite Harvestman Species from Croatia, with an Application of Confocal Laser Microscopy to Illustrate Genitalia in Opiliones". Breviora. 543 (1): 1–15. doi:10.3099/MCZ18.1. ISSN 0006-9698.
  5. Merino-Sáinz, Izaskun; Benavides, Ligia R.; Giribet, Gonzalo (2017-08-30). "The systematics and biogeography of the mite harvestman family Sironidae (Arachnida : Opiliones : Cyphophthalmi) with the description of five new species". Invertebrate Systematics. 31 (4): 456–491. doi:10.1071/IS16086. ISSN 1447-2600.
  6. Clouse, Ronald (2012). "The lineages of Stylocellidae (Arachnida: Opiliones: Cyphophthalmi)" (PDF). Zootaxa. 3595 (3595): 1–34. doi:10.11646/zootaxa.3595.1.1. ISSN 1175-5334.
  7. Kury, Adriano (2018). "The correct spelling of Leptopsalidinae (Arachnida: Opiliones: Cyphophthalmi)". Arachnida – Rivista Aracnologica Italiana. 18: 40–43.
  8. Clouse, Ronald; Schwendinger, Peter (2012). "Leptopsalis foveolata sp. n., a new species of Stylocellidae from Thailand that displays a novel morphological feature in the suborder Cyphophthalmi (Arachnida, Opiliones)" (PDF). Revue suisse de Zoologie. 119: 529–546. doi:10.5962/bhl.part.150204.
  9. Schmidt, Stephanie; Clouse, Ronald; Sharma, Prashant (2020). "A new Miopsalis from Mindanao supports a biogeographic umbilicus between Borneo and the southern Philippines (Arachnida: Opiliones: Cyphophthalmi: Stylocellidae)". Zootaxa. 4779 (3): 379–390. doi:10.11646/zootaxa.4779.3.6.
  10. Boyer, Sarah; Giribet, Gonzalo (2007). "A new model Gondwanan taxon: systematics and biogeography of the harvestman family Pettalidae (Arachnida, Opiliones, Cyphophthalmi), with a taxonomic revision of genera from Australia and New Zealand" (PDF). Cladistics. 23 (4): 337–361. doi:10.1111/j.1096-0031.2007.00149.x.
  11. Boyer, Sarah; Sharma, Prashant; Oberski, Jill; Coblens, Michelle; Popkin-Hall, Zachary; Jay, Katya (2016-04-05). "New species of Austropurcellia, cryptic short-range endemic mite harvestmen (Arachnida, Opiliones, Cyphophthalmi) from Australia's Wet Tropics biodiversity hotspot". ZooKeys (586): 37–93. doi:10.3897/zookeys.586.6774. ISSN 1313-2970. PMID 27199608.
  12. Giribet, Gonzalo (2003). "Karripurcellia, a new pettalid genus (Arachnida: Opiliones: Cyphophthalmi) from Western Australia, with a cladistic analysis of the family Pettalidae". Invertebrate Systematics. 17 (3): 387–406. doi:10.1071/IS02014.
  13. de Bivort, Benjamin; Giribet, Gonzalo (2010). "A systematic revision of the South African Pettalidae (Arachnida:Opiliones:Cyphophthalmi) based on a combined analysis of discrete and continuous morphological characters with the description of seven new species" (PDF). Invertebrate Systematics. 24 (4): 371–406. doi:10.1071/IS10015.
  14. Giribet, Gonzalo; Boyer, Sarah; Baker, Caitlin; et al. (2016). "A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) and the limits of taxonomic sampling" (PDF). Zoological Journal of the Linnean Society. 178 (3): 523–545. doi:10.1111/zoj.12419.
  15. Sharma, Prashant; Giribet, Gonzalo (2006). "A new Pettalus species (Opiliones, Cyphophthalmi, Pettalidae) from Sri Lanka with a discussion on the evolution of eyes in Cyphophthalmi" (PDF). The Journal of Arachnology. 34 (2): 331–341. doi:10.1636/H05-33.1.
  16. Sharma, Prashant; Karunarathna, Indika; Giribet, Gonzalo (2009). "On the endemic Sri Lankan genus Pettalus (Opiliones, Cyphophthalmi, Pettalidae) with a description of a new species and a discussion of its diversity" (PDF). Journal of Arachnology. 37 (1): 60–7. doi:10.1636/H08-56.1.
  17. Staręga, Wojciech (2008). "The second species of Purcellia Hansen et Sørensen, 1904 (Arachnida: Opiliones: Pettalidae) from South Africa" (PDF). Polish Journal of Entomology. 77 (1): 51–56.
  18. Benavides, Ligia; Hormiga, Gustavo; Giribet, Gonzalo (2019). "Phylogeny, evolution and systematic revision of the mite harvestman family Neogoveidae (Opiliones Cyphophthalmi)". Invertebrate Systematics. 33 (1): 101–180. doi:10.1071/IS18018.
  19. Clouse, Ronald; Wheeler, Ward (2014). "Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates". Zootaxa. 3814 (2): 177–201. doi:10.11646/zootaxa.3814.2.2. ISSN 1175-5334. PMID 24943422.
  20. Giribet, Gonzalo; Prieto, Carlos E. (2003-12-31). "A new Afrotropical Ogovea (Opiliones, Cyphophthalmi) from Cameroon, with a discussion on the taxonomic characters in the family Ogoveidae". Zootaxa. 329: 1–18. doi:10.5281/zenodo.156214.
  21. Sharma, Prashant; Giribet, Gonzalo (2009). "The family Troglosironidae (Opiliones: Cyphophthalmi) of New Caledonia". Zoologia Neocaledonica. 7. Mémoires du Muséum national d’Histoire naturelle. pp. 205–245. ISBN 9782856536186.
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