Doswell Formation

The Doswell Formation (also known as the Doswell Group) is a geologic unit of Upper Triassic age, part of the Newark Supergroup. The Doswell Formation was originally named to refer to a geological sequence which forms the lower part of the sedimentary fill of the Taylorsville Basin in Virginia and Maryland. This sequence was deposited by lakes and rivers in the developing rift basin.[1] However, a 2016 study determined that several geological layers in Pennsylvania as well as the neighboring Richmond Basin of Virginia also qualified as components of the Doswell Formation.[2]

Doswell Formation Stratigraphic range: Upper Triassic, Carnian
Type	Formation
Unit of	Chatham Group
Sub-units	Stagg Creek Member, Vinita Member, Irishtown Member, Lahaska Creek Member
Underlies	Stockton Formation
Overlies	Petersburg Granite, Evangeline Formation
Thickness	2,000 ft (610 m)
Location
Region	Virginia, Pennsylvania
Type section
Named for	Doswell, Virginia
Named by	Robert E. Weems

The most diverse and fossiliferous component of the Doswell formation is the Vinita member, also sometimes referred to as the Turkey Branch, Tuckahoe, or Falling Creek Formations in earlier publications. The Doswell formation is biostratigraphically characterized by a fauna including the fish Dictyopyge macrurus[3] and the conchostracan Laxitextella multireticulata.[4] The Richmond Basin has several notable fossil sites, such as the Tomahawk site which has hundreds of fossils from the cynodont Boreogomphodon,[5] and the Winterpock site which has an extraordinarily diverse assortment of plants.[3] The Taylorsville Basin is much more restricted in terms of fossil locales, but it was home to the unusual armored reptile Doswellia.[6]

The Doswell Formation is among the oldest Triassic formations on the east coast of the United States. It is believed to belong to the early Carnian (Cordevolian) age of the Triassic based on its fauna and flora,[4][2] which is distinctly dominated by tetrapods similar to gondwanan groups, as well as a high diversity of humidity-loving plants such as ferns and cycads. This gives it a distinct disconnect from the younger formations of the Newark Supergroup, which typically have a fauna similar to the Triassic formations of the western United States and a flora including elements such as conifer trees, which are better adapted for drier conditions.[3]

History and stratigraphy

Weems (1980)'s interpretation

The Doswell Formation was originally described by USGS paleontologist Robert E. Weems in 1980. He used it to refer to the entire Triassic geological sequence preserved at the Taylorsville basin, overlying the much older Carboniferous "Petersburg granite" which predated the basin, and underlying much younger Cretaceous and Cenozoic gravel. Weems subdivided the formation into several "members". The oldest of these (middle Carnian in age) was the Stagg Creek Member, a fluvial (river) deposit of sandstone and conglomerate.[7] Weems originally described this layer as lacking fossils, although later studies argued otherwise.[3]

Overlying the Stagg Creek Member was what Weems called the Falling Creek Member (late middle Carian). As sediments began to slow down the rivers of the Stagg Creek Member, lacustrine (lake) deposits began to form. Rivers still managed to flow into the now dominant lakes from different directions, depositing a diverse assortment of sediments. This allowed the Falling Creek Member to contain a variety of rock types, including sandstone, shale, siltstone, and occasionally even coal. In addition, it is the most fossiliferous part of the Taylorsville basin, containing the fossils of not only fish and invertebrates, but also reptiles such as the heavily armored Doswellia.[7]

The youngest (early late Carnian) and most geographically extensive member was the Newfound Member, named after the Newfound river. Outcrops belonging to this member either contain coarse sandstone and conglomerate, or much finer sandstone and siltstone. This member is believed to have been formed by a vast alluvial fan or delta created by southeastern-flowing rivers, and Araucarioxylon (petrified wood) is occasionally found within the member.[7]

Cornet & Olsen (1990)'s interpretation

In 1990, paleontologists Bruce Cornet & Paul Olsen described the Triassic fauna and flora of Virginia in detail. They argued that the Stagg Creek and Falling Creek Members actually coexisted at the same time, based on palynological correlations. In addition, they noted that the Stagg Creek actually preserved a few fossils, mainly of crustaceans and the abundant fish Dictyopyge.[3]

LeTourneau (2003)'s interpretation

In 2003, Columbia University geologist Peter LeTourneau became the first geologist to argue that the Doswell Formation was not the only Triassic geological layer in the Taylorsville Basin. He recognized the existence of a younger layer, which he called the King George Group. The King George Group was an extensive geological interval dominated by sandstone and conglomerate. LeTourneau also elevated the ranking of the Doswell Formation, renaming it to the Doswell Group. In conjunction with this, he also elevated the Falling Creek, and Newfound Members to formations. However, he also placed the Newfound Formation outside of the Doswell Group, instead placing it as the oldest unit of the King George Group.[1]

LeTourneau also evaluated Cornet & Olsen's claim that the Stagg Creek member coexisted with the Falling Creek Formation. He found that there were actually several distinct units grouped as part of the Stagg Creek unit. The original Stagg creek site which Weems (1980) based the member off of was found to belong to the middle portion of the Falling Creek Formation, therefore making the Stagg Creek Member part of that Formation. The portion of the Falling Creek Formation which was younger than the Stagg Creek Member was designated the Poor Farm Member, while the older portion was designated the Deer Creek Member. LeTourneau also added an additional formation to the Doswell Group, the South Anna Formation. This formation, the oldest section in the group, was very similar to the Stagg Creek Member in terms of its geological appearance and thickness, and as such South Anna outcroppings were originally considered to belong to the Stagg Creek Member according to Weems (1980). LeTournea differentiated the two based on their age, as determined by palynological dating.[1]

Weems, Tanner, & Lucas (2016)'s interpretation

In 2016, several paleontologists and geologists cooperated in a project which meant to correlate the individual Triassic basins of the Newark Supergroup with each other. They found that the Richmond and Taylorsville basins were likely deposited at the same time, a suspicion voiced earlier by Cornet & Olsen (1990).[3] In light of this revelation, they set out to link the formations and members of the Richmond Basin with those of the Taylorsville. They deranked the Doswell Group back to the Doswell Formation, and also found that the differences between the South Anna Formation and the Stagg Creek Member were not statistically significant. As a result, they abandoned the designation of the South Anna Formation, synonymizing it with the Stagg Creek Member as originally considered in 1980. They additionally found that the "barren beds" of the Richmond Basin (and several thinner slivers of strata in other basins) were also synonymous with the Stagg Creek Member.[2]

The Falling Creek Formation was more thoroughly deconstructed. Weems, Tanner, & Lucas found that this formation was basically identical to the Vinita Member (or "Vinita beds") of the Richmond basin. As the Vinita member was named approximately 70 years earlier than the Falling Creek Formation, it was considered to take priority in naming. The Poor Farm and Deer Creek members of the Falling Creek Formation were also abandoned due to being poorly defined by LeTourneau.[2]

These authors not only found the Doswell Formation to extend to the Richmond basin, but also to several other Triassic basins in Eastern Pennsylvania. For example, the "Irishtown beds" at the base of the Gettysburg basin were found to be a young layer of the Doswell Formation (the Irishtown Member) due to conchostracan dating. Lastly, an unusually old section of the Stockton Formation was also found to be a young part of the Doswell Formation (as the Lahaska Creek Member) due to preserving fossils of Calamops, a temnospondyl amphibian which lived at the same time as the Doswell Formation.[2]

Paleobiota

Tetrapods

Genus	Species	Sub-unit	Basin	Abundance	Notes	Images
Boreogomphodon[5]	B. jeffersoni	Vinita member (Tomahawk assemblage)	Richmond	"hundreds of specimens"	A traversodontid cynodont
Calamops[2]	C. paludosis	Lahaska Creek member	Newark		A temnospondyl
Doswellia[6]	D. kaltenbachi	Vinita member	Taylorsville, Richmond?	Partial skeleton, isolated bones	An archosauriform
Euscolosuchus[5]	E. olseni	Vinita member (Tomahawk assemblage)	Richmond	Armor scutes, vertebrae, and ribs	An archosauriform, possibly related to Crocodyliformes
Gomphiosauridion[8]	G. baileyae	Vinita member (Tomahawk assemblage)	Richmond	skull fragments	A procolophonid
Lacertilia indet.?[3]		Vinita member (Tomahawk assemblage)	Richmond
Lissamphibia indet.?[3]		Vinita member	Richmond	jaw fragment
Microconodon[5]	M. tenuirostris	Vinita member (Tomahawk assemblage)	Richmond	Teeth and jaw bones	A cynodont
Phytosauria indet.[5][6]		Vinita member	Richmond, Taylorsville	teeth
Rauisuchidae indet.[6]		Vinita member	Taylorsville?	limb bones
Rhynchocephalia indet.[5]		Vinita member (Tomahawk assemblage)	Richmond	skull fragments
Uatchitodon[5]	U. kroehleri	Vinita member (Tomahawk assemblage)	Richmond	teeth	A venomous reptile
Xenodiphyodon[8]	X. petraios	Vinita member (Tomahawk assemblage)	Richmond	partial jaw with teeth	A possible procolophonian

Fish

Genus	Species	Sub-unit	Basin	Abundance	Notes	Images
Cionichthys[3]	C. meekeri	Vinita member	Richmond, Taylorsville		A redfieldiid
Coelacanthiformes indet.[3]		Vinita member	Richmond
Dictyopyge[3]	D. macrurus	Stagg Creek member, Vinita member	Richmond, Taylorsville	Abundant	A redfieldiid
Lissodus	L. sp.	Vinita member (Tomahawk assemblage)	Richmond	teeth	A hybodont shark
Tanaocrossus[3]	T. sp.	Vinita member	Richmond		A palaeonisciform

Invertebrates

Genus	Species	Sub-unit	Basin	Abundance	Notes	Images
Darwinula[3]		Vinita member, Stagg Creek member	Richmond, Taylorsville		Ostracods
Gastropoda indet.[3]		Vinita member	Richmond, Taylorsville		Snails
Laxitextella[4]	L. multireticulata	Stagg Creek member, Vinita member, Irishtown member	Richmond, Taylorsville, Gettysburg	Abundant	A conchostracan (clam shrimp)
Unionidae indet.[3]		Vinita member	Taylorsville		Clams

Pteridophytes (spore-bearing plants)

[3]

Genus	Species	Sub-unit	Basin	Abundance	Notes	Images
Auriculophora	A. acrostichoides	Vinita member (Winterpock assemblage)	Richmond		A marattialean fern
Cladophlebis	C. auriculata, C. mexicana	Vinita member	Richmond		A filicalean fern
Clathropteris	C. meniscoides	Vinita member (Winterpock assemblage)	Richmond		A dipterid fern
Cyathocaulis	C. carolinensis	Vinita member (Winterpock assemblage)	Richmond		A cyatheaceaen tree fern
Cyathoforma (Asterotheca, Asterocarpus)	C. carolinensis, C. minuta, C. penticarpa	Stagg Creek member, Vinita member	Richmond, Taylorsville	common	A large tree fern
Danaeopsis	D. virginiensis	Vinita member (Winterpock assemblage)	Richmond		A marattialean fern
Dicranopteris	D. sp.	Vinita member (Winterpock assemblage)	Richmond		A forked fern
Equisetites	E. richmondensis, E. rogersii	Stagg Creek member, Vinita member	Richmond, Taylorsville	common	A horsetail
Gleichenites	G. distans	Vinita member (Winterpock assemblage)	Richmond	common	A forked fern
Isoetodendron	I. striata	Vinita member (Winterpock assemblage)	Richmond		A quillwort
Leptocyclotes	L. americana	Vinita member (Winterpock assemblage)	Richmond		A quillwort
Lonchopteris	L. oblonga	Vinita member (Winterpock assemblage)	Richmond		A fern
Mertensides	M. bullatus	Vinita member (Winterpock assemblage)	Richmond	common	A fern
Neocalamites	N. virginiensis,N. delawarensis	Stagg Creek member, Vinita member	Richmond, Taylorsville	common	A horsetail
Osmundites	O. winterpockensis	Vinita member (Winterpock assemblage)	Richmond		An osmunadcean fern
Pecopteris	P. rarinervis	Vinita member	Richmond		A type of leaf referable to several types of plants
Phlebopteris	P. smithii	Stagg Creek member	Taylorsville		A matoniacean fern
Pteridocaulis	P. rhombiformis, P. facialis	Vinita member (Winterpock assemblage)	Richmond		Stem of a tree fern
Todites (Acrostichites)	T. linnaeaefolius	Stagg Creek member, Vinita member	Richmond, Taylorsville	common	A fern

Gymnosperms

[3]

Genus	Species	Sub-unit	Basin	Abundance	Notes	Images
Macrotaniopteris	M. crassinervis, M. magnifolia	Stagg Creek member, Vinita member	Richmond, Taylorsville	common	A cycad
Plicarizamites	P. lanceolatus	Vinita member (Winterpock assemblage)	Richmond
Podozamites	P. lanceolatus, P. tenuistriatus	Stagg Creek member, Vinita member	Richmond, Taylorsville		A type of leaf referable to conifers
Primaraucaria	P. wielandii	Vinita member	Richmond	fairly common	An araucariacean conifer
Pterophyllum	P. affinae, P. braunianum, P. inaequale, P. giganteum, P. grandifolium, P. taxinum, P. tenuinervis	Stagg Creek member, Vinita member	Richmond, Taylorsville	Very common	A cycadeoid bennetitalean
Sagenopteris	S. rhoifolia	Vinita member (Winterpock assemblage)	Richmond		A seed fern
Sphenobaiera	S. striata	Stagg Creek member, Vinita member	Richmond, Taylorsville	common	A ginkgophyte
Sphenopteris	S. sitholeyi	Stagg Creek member, Vinita member	Richmond, Taylorsville	common	A seed fern
Sphenozamites	S. rogersianus	Stagg Creek member, Vinita member	Richmond, Taylorsville	common	A cycad
Stangerites	S. obliqua, S. planus	Vinita member (Winterpock assemblage)	Richmond		A cycad
Taeniopteris	T. diminuta	Vinita member (Winterpock assemblage)	Richmond	common	Leaves of a Pentoxylon-like plant
Triassiflorites	T. grandiflora	Vinita member (Winterpock assemblage)	Richmond		A cone, possibly from a bennetitalean
Zamiostrobus	Z. lissocardius	Vinita member (Winterpock assemblage)	Richmond		A cone from a cycad or conifer
Zamites	Z. powellii	Stagg Creek member	Taylorsville		A cycadeoid bennetitalean

References

1. LeTourneau, P.M. (2003). "Stratigraphic Architecture and Paleomagnetic Reversal Stratigraphy of the Late Triassic Taylorsville Basin, Virginia and Maryland". In LeTourneau P.M. & Olsen P.E. (ed.). The Great Rift Valleys of Pangea in Eastern North America: Volume 2. Edinburgh University Press. ISBN 978-0-231-12676-2. Retrieved 24 November 2012.
2. Weems, Robert E.; Tanner, Lawrence H.; Lucas, Spencer G. (2016). "Synthesis and revision of the lithostratigraphic groups and formations in the Upper Permian?–Lower Jurassic Newark Supergroup of eastern North America". Stratigraphy. 13 (2).
3. Cornet, Bruce; Olsen, Paul E. (1990). Early to Middle Carnian (Triassic) Flora and Fauna of the Richmond and Taylorsville Basins, Virginia and Maryland, U.S.A. Virginia Museum of Natural History. pp. 1–84. ISBN 0-9625801-1-2.
4. Kozur, Heinz W.; Weems, Robert E. (2010). "The biostratigraphic importance of conchostracans in the continental Triassic of the northern hemisphere" (PDF). Geological Society of London, Special Publications. 334 (1): 315–417. Bibcode:2010GSLSP.334..315K. doi:10.1144/SP334.13. S2CID 131224365.
5. Sues, Hans-Dieter; Olson, Paul E.; Kroehler, Peter A. (1994). Small tetrapods from the Upper Triassic of the Richmond basin (Newark Supergroup), Virginia (PDF). In the Shadow of the Dinosaurs. Cambridge University Press. pp. 161–170.
6. R. E. Weems (1980). "An unusual newly discovered archosaur from the Upper Triassic of Virginia, U.S.A." Transactions of the American Philosophical Society. New Series. 70 (7): 1–53. doi:10.2307/1006472. JSTOR 1006472.
7. Weems, Robert E. (1980). Geology of the Taylorsville basin, Hanover County, Virginia (PDF). Contributions to Virginia Geology- IV. Department of Conservation and Economic Development. pp. 23–38.
8. Sues, Hans-Dieter; Olson, Paul E. (1992). "A new procolophonid and a new tetrapod of uncertain, possibly procolophonian affinities from the Upper Triassic of Virginia" (PDF). Journal of Vertebrate Paleontology. 13 (3): 282–286. doi:10.1080/02724634.1993.10011510.