Escucha Formation

The Escucha Formation is a geological formation in La Rioja and Teruel provinces of northeastern Spain whose strata date back to the late Aptian to middle Albian stages of the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.[1]

Escucha Formation
Stratigraphic range: Late Aptian-mid Albian
~117–103 Ma
View of Escucha and surrounding rocks
TypeGeological formation
Sub-unitsE1 to E3
UnderliesUtrillas Formation
OverliesCastrillo de la Reina, Benassal & Oliete Formations
Thickness~70 m (230 ft)
Lithology
PrimarySiltstone, mudstone (E1, E3) sandstone (E2)
OtherCoal (E1) siltstone (E2), amber
Location
Coordinates42.6°N 2.7°W / 42.6; -2.7
Approximate paleocoordinates30.4°N 1.4°E / 30.4; 1.4
RegionTeruel, Aragón
La Rioja
Country Spain
ExtentCameros, Maestrazgo & Oliete Basins
Type section
Named forEscucha, Teruel
Escucha Formation (Spain)

The approximately 70 m (230 ft) thick formation underlies the Utrillas Formation and overlies Castrillo de la Reina, Benassal & Oliete Formations. The Escucha Formation comprises siltstones, mudstones, sandstones, coal, siltstones and amber, in which several fossil insects were found. The formation was deposited in a variety of continental to paralic (deltaic environments.[2]

Fossil content

The Escucha Formation has provided the following fossils, among others:[3]

Dinosaurs

Dinosaurs
Taxa Location Notes Images
Europelta carbonensis La Rioja [4]
Proa valdearinnoensis La Rioja [5]
Other dinosaurs
Reptiles
Fish

Insects

Insects
Taxa Location/site Notes Images
Alavaraphidia Peñacerrada I amber [7]
Amarantoraphidia Peñacerrada I amber [7]
Microcostaphron parvus Peñacerrada I amber [8]
Radiophron ibericus Peñacerrada I amber [9]
Styporaphidia? hispanica Peñacerrada I amber [7]
Other arthropods
  • Alavia neli
  • Archaelagonops alavensis
  • Mesozygiella dunlopi
  • Alavesiaphis margaritae
  • Hispanocader lisae
  • Alavametra popovi
  • Iberovelia quisquilia
  • Glaesivelia pulcherrima
  • Gymnopollisthrips maior
  • G. minor
  • Manicapsocidus enigmaticus
  • Archaeatropos alavensis
  • Preempheria antiqua
  • Empheropsocus arilloi
  • E. margineglabrus
  • Morazatermes krishnai
  • Cantabritermes simplex
  • Autrigonoforceps iberica
  • Hispanelcana alavensis
  • H. arilloi
  • H. lopezvallei
  • Libanophron sugaar
  • Hippocoon basajauni
  • Burmaphron iratxoak
  • B. jentilak
  • B. sorginak
  • Tagsmiphron olentzero
  • Elasmophron mari
  • Ectenobythus iberiensis
  • Liztor pilosus
  • Cretepyris martini
  • Zophepyris alavaensis
  • Ampulicomorpha perialla
  • Microcostaphron parvus
  • Archephedrus stolamissus
  • Protorhyssalopsis perrichoti
  • Spathiopteryx alavarommopsis
  • Serphites lamiak
  • Aposerphites angustus
  • Archaeromma hispanicum
  • Galloromma alavaensis
  • Alavaromma orchamum
  • Tithonoscelio resinalis
  • Bruescelio platycephalus
  • Electroteleiopsis hebdomas
  • Alavascelio delvallei
  • Amissascelio temporarius
  • Perimoscelio tyrbastes
  • Perimoscelio confector
  • Proterosceliopsis masneri
  • Juxtascelio interitus
  • Iberopria perialla
  • Iberoevania roblesi
  • Cretevania alonsoi
  • Iberomaimetsha nihtmara
  • Iberomaimetsha rasnitsyni
  • Valaa delclosi
  • Megalava truncata
  • Eosyntexis parva
  • Cretasonoma corinformibus
  • Penarhytus tenebris
  • Prosolierius parvus
  • Cretakarenni hispanicus
  • Rhizophtoma longus
  • Darwinylus marcosi
  • Mediumiuga sinespinis
  • Archiaustroconops alavensis
  • Leptoconops (Leptoconops) zherikhini
  • Gerontodacus skalskii
  • Chimeromyia alava
  • Chimeromyina concilia
  • Euliphora grimaldii
  • Alavesia subiasi
  • Tethepomima holomma
  • Tethepomyia buruhandi
  • Lysistrata emerita
  • Hegalari minor
  • Hegalari antzinako
  • Alavamanota hispanica
  • Allocotocera xavieri
  • Cretohaplusia ortunoi
  • Eltxo cretaceus
  • Helius (Helius) alavensis
  • Helius (Helius) spiralis
  • Espanoderus barbarae
  • Alavaraphidia imperterrita
  • Baissoptera cretaceoelectra
  • Cretokatianna bucculenta
  • Sphyrotheciscus senectus
  • Archeallacma dolichopoda
  • Katiannasminthurus xenopygus
  • Pseudosminthurides stoechus
  • Burmisotoma spinulifera
  • Protoisotoma autrigoniensis
  • Proisotoma communis
  • Proleptochelia tenuissima
  • Electrotanais monolithus
  • Alavatanais carabe
  • Alavatanais margulisae
  • Eurotanais terminator
  • Spinomegops arcanus
  • Strieremaeus minguezae
  • Iberofoveopsis miguelesi
  • Hispanothrips utrillensis
  • Aragonitermes teruelensis
  • Aragonimantis aenigma
  • Mymaropsis turolensis
  • Serphites silban
  • Galloromma turolensis
  • Alavaromma orchamum
  • Cretevania rubusensis
  • Cretevania alcalai
  • Cretevania montoyai
  • Actenobius magneoculus
  • Arra legalovi
  • Leptoconops (Leptoconops) zherikhini
  • Atriculicoides sanjusti
  • Atriculicoides hispanicus
  • Gerontodacus skalskii
  • Microphorites utrillensis
  • Litoleptis fossilis
  • Burmazelmira grimaldii
  • Aragomantispa lacerata
  • Spinomegops aragonensis
  • Ametroproctus valeriae
  • Cretaceobodes martinezae
  • Trhypochthonius lopezvallei
Invertebrates
Flora

Correlation

Early Cretaceous stratigraphy of Iberia
MaAgePaleomap \ BasinsCantabrianOlanyàCamerosMaestrazgoOlieteGalveMorellaSouth IberianPre-beticLusitanian
100Cenomanian
La CabanaSopeiraUtrillasMosquerelaCaranguejeira
AltamiraUtrillas
Eguino
125AlbianUllaga - BalmasedaLluçàTraiguera
Monte GrandeEscuchaEscuchaJijona
Itxina - Miono
AptianValmaseda - TellamendiOl Gp. - CastrilloBenassalBenassalOlhos
FontEn Gp. - LezaMorella/OlieteOlieteVillaroyaMorellaCapas
Rojas
Almargem
Patrocinio - ErnagaSenyúsEn Gp. - JubelaForcallVillaroyaUpper
Bedoulian
Figueira
BarremianVega de PasCabóAbejarXertAlacónXertHuérguinaAssises
PradaArtolesColladoMoutonianum Papo Seco
RúbiesTera Gp. - GolmayoAlacón/BlesaBlesaCamarillasMirambel
150HauterivianUr Gp. - PinillaLlacovaCastellarTera Gp. - PinillaVillaresPorto da
Calada
hiatus
HuervaGaita
ValanginianVillaroUr Gp. - LarribaPed Gp. - Hortigüela
Ped Gp. - HortigüelaPed Gp. - Piedrahita
PeñacobaGalveMiravetes
BerriasianCab Gp. - ArceraValdepradohiatusAlfambra
TdL Gp. - RupeloArzobispohiatusTollo
On Gp. - Huérteles
Sierra Matute
TithonianLastresTera Gp. - MagañaHiguerelesTera Gp. - MagañaLourinhã
Arzobispo
Ágreda
Legend Major fossiliferous, oofossiliferous, ichnofossiliferous, coproliferous, minor formation
Sources

See also

References

  1. Weishampel, et al., 2004
  2. Peyrot et al., 2007
  3. Escucha Formation at Fossilworks.org
  4. Kirkland et al., 2013
  5. McDonald et al., 2012
  6. Pérez García et al., 2020
  7. Pérez de la Fuente et al., 2012
  8. Ortega Blanco et al., 2010, p.272
  9. Ortega Blanco et al., 2010, p.267

Bibliography

  • Kirkland, J.I.; L. Alcalá; M.A. Loewen; E. Espílez; L. Mampel, and J.P. Wiersma. 2013. The Basal Nodosaurid Ankylosaur Europelta carbonensis n. gen., n. sp. From the Lower Cretaceous (Lower Albian) Escucha Formation of Northeastern Spain. PLoS ONE 8. e80405. Accessed 2019-10-09.
  • Ortega Blanco, J.; A.P. Rasnitsyn, and X. Delclòs. 2010. A new family of ceraphronoid wasps from Early Cretaceous Álava Amber, Spain. Acta Palaeontologica Polonica 55. 265–276. Accessed 2019-10-09.
  • Pérez de la Fuente, R.; E. Peñalver; X. Delclòs, and M.S. Engel. 2012. Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida, Raphidioptera). ZooKeys 204. 1–40. doi:10.3897/zookeys.204.2740 PMID 22787417 PMC 3391719
  • Pérez García, A.; E. Espílez; L. Mampel, and L. Alcalá. 2020. A new basal turtle represented by the two most complete skeletons of Helochelydridae in Europe. Cretaceous Research 107. Article 104291. doi:10.1016/j.cretres.2019.104291
  • Peyrot, Daniel; Juan Pedro Rodríguez López; Luis Lassaletta; Nieves Meléndez, and Eduardo Barrón. 2007. Contributions to the palaeoenvironmental knowledge of the Escucha Formation in the Lower Cretaceous Oliete Sub-basin, Teruel, Spain. Comptes Rendus Palevol 6. 469–481. doi:10.1016/j.crpv.2007.09.014ISSN 1631-0683
  • Weishampel, David B.; Peter Dodson, and Halszka Osmólska (eds.). 2004. The Dinosauria, 2nd edition, 1–880. Berkeley: University of California Press. Accessed 2019-02-21.ISBN 0-520-24209-2

Further reading

  • L. Alcalá, E. Espílez, L. Mampel, J. I. Kirkland, M. Ortiga, D. Rubio, A. González, D. Ayala, A. Cobos, R. Royo-Torres, F. Gascó and M. D. Pesquero. 2012. A new Lower Cretaceous vertebrate bonebed near Arino (Tereul, Aragon, Spain); found and managed in a joint collaboration between a mining company and a palaeontological park. Geoheritage 4:275-286
  • J. I. Canudo, A. Cobos, C. Martín-Closas, X. Murelaga, X. Pereda-Suberbiola, R. Royo-Torres, J. I. Ruiz-Omeñaca and L. M. Sender. 2005. Sobre la presencia de dinosaurios ornitópodos en la Formación Escucha (Cretácico Inferior, Albiense): redescubierto “Iguanodon” en Utrillas (Teruel) [On the presence of ornithopod dinosaurs in the Escucha Formation (Lower Cretaceous, Albian): redescribing “Iguanodon” in Utrillas (Teruel)]. Fundamental 6:51-56
  • E. Peñalver, D. A. Grimaldi, and X. Delclos. 2006. Early Cretaceous spider web with its prey. Science 312:1761
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