Hemicyoninae

Hemicyoninae is an extinct subfamily of Ursidae[2][1] often called "dog-bears", literally "half dog" (Greek: ἡμικυων hemi-kyōn), bear-like carnivoran living in Europe, North America, Africa and Asia during the Oligocene through Miocene epochs 33.9–5.3 Ma, existing for approximately 28.6 million years . They are sometimes classified as a separate family.[3]

Hemicyoninae Temporal range: Early Oligocene to Late Miocene 33.9–5.3 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Hemicyon sansaniensis
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Ursidae
Subfamily:	†Hemicyoninae Frick, 1926
Tribes and genera
Tribe †Cephalogalini †Adelpharctos[1] †Cyonarctos[2] †Phoberogale[2] †Filholictis[2] †Cephalogale[2] Tribe †Phoberocyonini †Plithocyon †Phoberocyon Tribe †Hemicyonini †Zaragocyon †Dinocyon †Hemicyon

Systematics

• Subfamily †Hemicyoninae (Frick, 1926)
  • Tribe †Cephalogalini (de Bonis, 2013)
    • †Adelpharctos (de Bonis, 1971)
      • †Adelpharctos ginsburgi (de Bonis, 2011)
      • †Adelpharctos mirus (de Bonis, 1971)
    • †Cyonarctos (de Bonis, 2013)
      • †Cyonarctos dessei (de Bonis, 2013)
    • †Phoberogale (Ginsburg & Morales, 1995)
      • †Phoberogale minor (Filhol, 1877)
      • †Phoberogale bonali (Helbing, 1928)
      • †Phoberogale depereti (Viret, 1929)
      • †Phoberogale gracile (Pomel, 1847)
    • †Filholictis (de Bonis, 2013)
      • †Filholictis filholi (Munier-Chalmas, 1877)
    • †Cephalogale (Jourdan, 1862)
      • †Cephalogale shareri (Wang, et al., 2009)
      • †Cephalogale gergoviensis (Viret, 1929)
      • †Cephalogale ginesticus (Kuss, 1962)
      • †Cephalogale geoffroyi (Jourdan, 1862)
  • Tribe †Phoberocyonini (Ginsburg & Morales, 1995)
    • †Plithocyon (Ginsburg, 1955)
      • †Plithocyon armagnacensis (Ginsburg, 1955)
      • †Plithocyon statzlingii (Frick, 1926)
      • †Plithocyon bruneti (Ginsburg, 1980)
      • †Plithocyon barstowensis (Frick, 1926)
      • †Plithocyon ursinus (Cope, 1875)
    • †Phoberocyon (Ginsburg, 1955)
      • †Phoberocyon hispanicus (Ginsburg & Morales, 1998)
      • †Phoberocyon dehmi (Ginsburg, 1955)
      • †Phoberocyon huerzeleri (Ginsburg, 1955)
      • †Phoberocyon aurelianensis (Mayet, 1908)
      • †Phoberocyon youngi (Xiang et al., 1986)
      • †Phoberocyon johnhenryi (White, 1947)
  • Tribe †Hemicyonini (Frick, 1926)
    • †Zaragocyon (Ginsburg & Morales, 1995)
      • †Zaragocyon daamsi (Ginsburg & Morales, 1995)
    • †Dinocyon (Jourdan, 1861)
      • †Dinocyon aurelianensis (Frick, 1926)
      • †Dinocyon sansaniensis (Frick, 1926)
      • †Dinocyon thenardi (Jourdan, 1861)
    • †Hemicyon (Lartet, 1851)
      • †Hemicyon barbouri (Colbert, 1941)
      • †Hemicyon teilhardi (Colbert, 1939)
      • †Hemicyon grivensis (Frick, 1926)
      • †Hemicyon minor (Dépéret, 1887)
      • †Hemicyon sansaniensis (Lartet, 1851)

References

1. L. de Bonis (2011). "A new species of Adelpharctos (Mammalia, Carnivora, Ursidae) from the late Oligocene of the "Phosphorites du Quercy" (France)". Estudios Geológicos. 67 (2): 179–186. doi:10.3989/egeol.40553.181.
2. Louis De Bonis (2013). "Ursidae (Mammalia, Carnivora) from the Late Oligocene of the "Phosphorites du Quercy" (France) and a reappraisal of the genus Cephalogale Geoffroy, 1862". Geodiversitas. 35 (4): 787–814. doi:10.5252/g2013n4a4.
3. McKenna, Malcolm C., and Bell, Susan K. 1997. Classification of Mammals Above the Species Level. Columbia University Press, New York, 631 pp. ISBN 0-231-11013-8