Isatis

Isatis is a genus of flowering plants in the family Brassicaceae, native to the Mediterranean region east to central Asia. The genus includes woad (Isatis tinctoria). Due to their extremely variable morphology, the Asian species in particular are difficult to determine; the only reliable diagnostic feature is the ripe fruit. They are (usually) biennial or perennial herbaceous plants, often bluish and hairless or downy hairy with the upright stem branched.[2]

Isatis
Isatis tinctoria
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Brassicales
Family: Brassicaceae
Genus: Isatis
L.
Species

About 30 species, including:
Isatis boissieriana
 Isatis glauca
Isatis tinctoria
others (see text)

Synonyms[1]

Description

They are annual, biennial or perennial, branched herbs, usually glabrous and glaucous except silicon. Basal leaves generally elliptic-oblong, sessile; sessile caulinary, rounded to oval-oblong.

The hermaphrodite flowers are fourfold double perianth. The four sepals are ascending to upright. The four yellow to off-white or lilac-white petals are at least as long as the sepals. They have six stamens with very small, egg-shaped or elongated-round anthers. There are nectar glands. Racemose is branched or paniculated, ebracted, inflorescence, often reaching lax and elongated in the fruit.

The fruit is a generally linear silicon, oblong-cuneate to suborbicular, indehiscent, flattened laterally, unilocular, little to conspicuously winged, glabrous or with tiny hairs.[3]

Species

Currently accepted species include:[1]

  • Isatis afghanica Hadac & Chrtek
  • Isatis amani P.H.Davis
  • Isatis apennina Ten. ex Grande
  • Isatis apscheronica N.Busch
  • Isatis aptera (Boiss. & Heldr.) Al-Shehbaz, Moazzeni & Mumm.
  • Isatis arenaria Azn.
  • Isatis armena L.
  • Isatis arnoldiana N.Busch
  • Isatis aucheri Boiss.
  • Isatis biscutellifolia Boiss. & Buhse
  • Isatis bitlisica P.H.Davis
  • Isatis boissieriana Rchb.f.
  • Isatis brachycarpa C.A.Mey.
  • Isatis brevipes (Bunge) Jafri
  • Isatis bullata Aitch. & Hemsl.
  • Isatis bungeana Seidlitz
  • Isatis buschiana Schischk.
  • Isatis callifera Boiss. & Balansa
  • Isatis campylocarpa Boiss.
  • Isatis canaliculata (Vassilcz.) V.V.Botschantz.
  • Isatis candolleana Boiss.
  • Isatis cappadocica Desv.
  • Isatis cardiocarpa (Trautv.) Al-Shehbaz, Moazzeni & Mumm.
  • Isatis caucasica N.Busch
  • Isatis cochlearis Boiss.
  • Isatis constricta P.H.Davis
  • Isatis costata C.A.Mey.
  • Isatis davisiana H.Misirdali ex P.H.Davis, R.R.Mill & Kit Tan
  • Isatis demiriziana H.Misirdali ex P.H.Davis, R.R.Mill & Kit Tan
  • Isatis densiflora (Bunge ex Boiss.) D.A.German
  • Isatis deserti (N.Busch) V.V.Botschantz.
  • Isatis djurjaedae Coss. & Durieu
  • Isatis elegans (Boiss.) Hadac & Chrtek
  • Isatis emarginata Kar. & Kir.
  • Isatis erzurumica P.H.Davis
  • Isatis floribunda Boiss. ex Bornm.
  • Isatis frigida Boiss. & Kotschy
  • Isatis frutescens Kar. & Kir.
  • Isatis gaubae Bornm.
  • Isatis glastifolia (Fisch. & C.A.Mey.) Al-Shehbaz, Moazzeni & Mumm.
  • Isatis glauca Aucher ex Boiss.
  • Isatis grammotis Kit Tan
  • Isatis gymnocarpa (Fisch. ex DC.) Al-Shehbaz, Moazzeni & Mumm.
  • Isatis harsukhii O.E.Schulz
  • Isatis hirtocalyx Franch.
  • Isatis huber-morathii P.H.Davis
  • Isatis iberica Steven
  • Isatis jacutensis (N.Busch) N.Busch
  • Isatis karjaginii Schischk.
  • Isatis kotschyana Boiss. & Hohen.
  • Isatis kozlovskyi Grossh.
  • Isatis laevigata Trautv.
  • Isatis latisiliqua Steven
  • Isatis leuconeura Boiss. & Buhse
  • Isatis littoralis Steven
  • Isatis lockmanniana Kotschy ex Boiss.
  • Isatis lusitanica L.
  • Isatis mardinensis P.H.Davis & H.Misirdali
  • Isatis maxima Pavlov
  • Isatis microcarpa J.Gay ex Boiss.
  • Isatis minima Bunge
  • Isatis multicaulis (Kar. & Kir.) Jafri
  • Isatis oblongata DC.
  • Isatis odontogera (Bordz.) D.A.German
  • Isatis ornithorhynchus N.Busch
  • Isatis pachycarpa Rech.f., Aellen & Esfand.
  • Isatis pinnatiloba P.H.Davis
  • Isatis platyloba Link ex Steud.
  • Isatis praecox Kit. ex Tratt.
  • Isatis raimondoi Di Grist., Scafidi & Domina
  • Isatis raphanifolia Boiss.
  • Isatis rugulosa Bunge ex Boiss.
  • Isatis sabulosa Steven ex Ledeb.
  • Isatis sevangensis N.Busch
  • Isatis sivasica P.H.Davis
  • Isatis spatella P.H.Davis
  • Isatis spectabilis P.H.Davis
  • Isatis steveniana Trautv.
  • Isatis stocksii Boiss.
  • Isatis subdidyma (N.Busch) V.E.Avet.
  • Isatis takhtajanii Avet.
  • Isatis tinctoria L.
  • Isatis tomentella Boiss. & Balansa
  • Isatis trachycarpa Trautv.
  • Isatis turcomanica Korsh.
  • Isatis undulata Aucher ex Boiss.
  • Isatis violascens Bunge
  • Isatis zarrei Al-Shehbaz, Moazzeni & Mumm.

References
  1. "Isatis Tourn. ex L." Plants of the World Online. Board of Trustees of the Royal Botanic Gardens, Kew. 2017. Retrieved 13 August 2020.
  2. H. Moazzeni et al .: Phylogeny of Isatis (Brassicaceae) and allied genera based on ITS sequences of nuclear ribosomal DNA and morphological characters. In: Flora (Elsevier) , Volume 205, 2010, pp. 337–343.
  3. Flora of North America Editorial Committee, e. 2010. Magnoliophyta: Salicaceae to Brassicaceae. Fl. N. Amer. 7: i – xxii, 1–797.


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