Jeholosauridae

Jeholosaurids were herbivorous neornithischian dinosaurs from the Cretaceous Period (Aptian - Santonian, with a possible Campanian record) of Asia. The family was first proposed by Han et al. in 2012. The jeholosaurids were defined as those ornithischians more closely related to Jeholosaurus shangyuanensis than to Hypsilophodon foxii, Iguanodon bernissartensis, Protoceratops andrewsi, Pachycephalosaurus wyomingensis, or Thescelosaurus neglectus. The Jeholosauridae includes the type genus Jeholosaurus, and also possibly Yueosaurus, Changchunsaurus, and Haya.[1][2][3]

Jeholosauridae Temporal range: Early to Late Cretaceous, PreꞒ Ꞓ O S D C P T J K Pg N Possible Campanian record
Jeholosaurus shangyuanensis skeleton reconstruction (Dinosaurland, Lyme Regis).
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Order:	†Ornithischia
Clade:	†Neornithischia
Family:	†Jeholosauridae Han et al., 2012
Genera
†Jeholosaurus (type genus) †Yueosaurus? †Changchunsaurus? †Haya?

The cladogram below results from analysis by Herne et al., 2019, which added Changchunsaurus and Haya to Jeholosauridae, but moved Yueosaurus to the sister family Thescelosauridae.[3]

Ornithischia

Heterodontosauridae       Eocursor       Thyreophora   Neornithischia   Lesothosaurus       Agilisaurus         Hexinlusaurus     Yandusaurus           Nanosaurus     Jeholosauridae   Haya       Jeholosaurus     Changchunsaurus       Thescelosauridae     Orodromeus     Koreanosaurus         Zephyrosaurus       Yueosaurus     Thescelosaurus             Cerapoda   Marginocephalia   Ornithopoda   Parksosaurus     Elasmaria   Talenkauen         Macrogryphosaurus     Gasparinisaura         Galleonosaurus     Leaellynasaura     Anabisetia     Diluvicursor         Clypeodonta   Hypsilophodon   Iguanodontia   Rhabdodontidae       Muttaburrasaurus       Tenontosaurus     Dryomorpha

References

1. Han, Feng-Lu; Paul M. Barrett; Richard J. Butler; Xing Xu (2012). "Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China". Journal of Vertebrate Paleontology. 32 (6): 1370–1395. doi:10.1080/02724634.2012.694385.
2. Boyd CA. (2015) The systematic relationships and biogeographic history of ornithischian dinosaurs. PeerJ 3:e1523 https://doi.org/10.7717/peerj.1523
3. Herne, Matthew C.; Nair, Jay P.; Evans, Alistair R.; Tait, Alan M. (2019). "New small-bodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian-Antarctic rift system, with revision of Qantassaurus intrepidus Rich and Vickers-Rich, 1999". Journal of Paleontology. doi:10.1017/jpa.2018.95.