Sceptridium

Sceptridium is a genus of seedless vascular plants in the family Ophioglossaceae,[1] closely allied to (and often included as a subgenus[2] of) the genus Botrychium (the moonworts and grapeferns). It is also closely related to the genus Botrypus (the rattlesnake fern, often treated as the subgenus Osmundopteris under Botrychium). Sceptridium species are commonly called the grape-ferns.

Sceptridium
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Class: Polypodiopsida
Order: Ophioglossales
Family: Ophioglossaceae
Subfamily: Botrychioideae
Genus: Sceptridium
Lyon 1905
Species

See text

These plants are small with fleshy roots, and reproduce by spores shed into the air. They differ from the moonworts in having at least some sterile fronds (all fronds in Botrychium are spore-bearing), and in the fronds being bi- or tri-pinnate (Botrychium are single pinnate, or rarely bipinnate); and from Botrypus in being evergreen, or at least winter-green (Botrypus are deciduous) and having the non-spore-bearing part of the frond long-stalked (short-stalked in Botrypus).

Species

Sceptridium species and their Botrychium synonyms include:

  • Sceptridium dissectum (Spreng.) Lyon 1905 – Cut-leaved grape-fern[3]
    • Botrychium dissectum Spreng. 1804
    • Botrychium obliquum Muhl. 1810[4]
  • Sceptridium jenmanii (Underw.) Lyon 1905 – Alabama grape-fern
    • Botrychium jenmanii Underw. 1900
    • Botrychium alabamense Maxon 1906[5]
  • Sceptridium multifidum (S.G.Gmel.) Nishida ex Tagawa 1958 – Leather grape-fern[3]
    • Osmunda multifida S.G. Gmel. 1768
    • Botrychium silaifolium C.Presl 1825
    • Botrychium multifidum (S.G. Gmel.) Rupr. 1859
    • Botrychium coulteri Underw. 1898
    • Botrychium californicum Underw. 1905[6]
  • Sceptridium lunarioides (Michx.) Holub - Winter grapefern.[7]
    • Botrychium lunarioides (Michx.) Sw.[8]
  • Sceptridium oneidense (Gilbert) Holub 1998 – Blunt-leaved grape-fern[3] (northeastern North America)[9]
    • Botrychium oneidense (Gilbert) House 1905[10]
  • Sceptridium rugulosum (W.H.Wagner) Skoda & Holub 1996 – Ternate grape-fern[3]
    • Botrychium rugulosum W.H.Wagner 1982[11]
  • Sceptridium subbifoliatum (Brack.) Lyon 1905 – Island grape-fern[12]
    • Botrychium subbifoliatum Brack. 1854[13]

References
  1. Christenhusz, Maarten J. M.; Zhang, Xian-Chun; Schneider, Harald (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19: 7–54. doi:10.11646/phytotaxa.19.1.2.
  2. subgenus Tropicos.org. Missouri Botanical Garden. 16 Jan 2012
  3. MICHIGAN FLORA ONLINE. A. A. Reznicek, E. G. Voss, & B. S. Walters. February 2011. University of Michigan. Web. January 16, 2012
  4. Botrychium obliquum Tropicos.org. Missouri Botanical Garden. 16 Jan 2012
  5. Botrychium alabamense Tropicos.org. Missouri Botanical Garden. 16 Jan 2012
  6. Botrychium multifidum Tropicos.org. Missouri Botanical Garden. 16 Jan 2012
  7. "PteridoPortal - Sceptridium lunarioides". www.pteridoportal.org. Retrieved 7 October 2020.
  8. "NatureServe Explorer 2.0 - Botrychium lunarioides Winter Grapefern". explorer.natureserve.org. Retrieved 7 October 2020.
  9. "Botrychium oneidense". Natural Resources Conservation Service PLANTS Database. USDA. Retrieved 16 Jan 2012.
  10. Botrychium oneidense Tropicos.org. Missouri Botanical Garden. 16 Jan 2012
  11. Botrychium rugulosum Tropicos.org. Missouri Botanical Garden. 16 Jan 2012
  12. "Sceptridium subbifoliatum". Natural Resources Conservation Service PLANTS Database. USDA. Retrieved 16 Jan 2012.
  13. Botrychium subbifoliatum International Plant Names Index (IPNI) 16 Jan 2012


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