Titanis

Titanis is an extinct genus of giant flightless terror birds that inhabited North America during the early Pliocene to early Pleistocene epochs.[1][2] The generic name, Titanis, refers to the titans, Ancient Greek gods that preceded the Twelve Olympians, in allusion to the bird's size. The specific name, T. walleri, honors the holotype's collector, Benjamin I. Waller,[3] an avocational underwater archaeologist.[4][5] Titanis was thought to be carnivorous and most likely preyed on the many small mammals of the time period.[6] This giant flightless bird was one of the most efficient predators of its time in North America.[7]

Titanis
Temporal range: Early Pliocene to Early Pleistocene, 5–1.8 Ma
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Reconstructed skeleton, Florida Museum of Natural History
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Cariamiformes
Family: †Phorusrhacidae
Subfamily: †Phorusrhacinae
Genus: †Titanis
Brodkorb, 1963
Type species
†Titanis walleri
Brodkorb, 1963

Description
Life reconstruction

Titanis was 2.5 m (8 ft 2 in) tall and weighed approximately 150 kilograms (330 lb).[7][8] When compared with other phorusrhacids, the examined material indicates a large variation in the size of Titanis, perhaps indicating strong sexual dimorphism.[9] It had long, agile legs, and three-toed feet with long talons. It could undoubtedly run at high speeds when hunting.[7] Though its skull has not been found, it most probably would have been large, with a huge, axe-like beak, like its relatives.

Overall, Titanis was very similar to the South American Phorusrhacos and Devincenzia, its closest relatives. However, it differs from these in having a shorter, thicker neck, and an overall more heavily built bodily structure. Little is known of its body structure, but it seems to have been less wide-footed than Devincenzia, with a proportionally much stronger middle toe.[10]

Wings

The wings were small and could not have been used for flight. The wing bones articulated in an unusual joint-like structure, suggesting the digits could flex to some degree. It also had a relatively rigid wrist, which would not have allowed the hand to fold back against the arm to the same degree as other birds. This led R. M. Chandler to suggest that the wings may have supported some type of clawed, mobile hand similar to the hands of non-avian theropod dinosaurs, such as the dromaeosaurs (also popularly known as "raptors").[11]

However, it was later pointed out that this wing joint is not in fact unique, and is present in seriemas (extant members of the same order, Cariamae, to which Titanis and other phorusrhacids belonged), which do not have any specialized grasping hands.[12]

Discovery
Pierce Brodkorb with the tarsometatarsus of Titanis (dark) and another bird

It lived approximately 5 to 2 million years ago (early Pliocene to early Pleistocene) in North America. Fossil evidence has been found in three locations in Florida and one in Texas.[13] The Gilchrist County, Florida site dates from 3.0 to 2.9 million years ago.[14] The Santa Fe River, where 27 of the 41 Titanis fossil specimen have been found, is located in Gilchrist County. The other locations that Titanis has been found include Port Charlotte and Inglis, Florida. Only one specimen has been found outside of Florida, and that was in the Nueces River in Texas.

Classification

Titanis was part of the group of giant flightless birds called the Phorusrhacidae, which are nicknamed "terror birds". It was thought to represent the youngest species of the lineage but recently, a significantly younger South American example has been reported.[15] The Phorusrhacidae originated in South America; Titanis is the only known member of the branch of the group that migrated out of that continent during the Great American Interchange.

A lineage of related predatory birds, the bathornithids, occurred in North America from the Paleocene to the Miocene. They were not ancestral to Titanis or any other phorusrhacid, but they occupied similar ecological niches and some like Paracrax even attained similar sizes, reaching above 2 meters in height. They became extinct more or less 15 million years before Titanis reached North America.[16] The only living relatives of Titanis are the seriemas.

Paleobiology

Studies of the closely related Andalgalornis steulleti, which is also in the family Phorusrhacidae reveals new information about the head and neck movement of T. walleri and all "terror birds" of this family. In a series of tests on the skull of Andalgalornis, it was found that these birds would have had a hard time moving their heads laterally. However, the back and forth movement of the skull was tremendous, and it is thought that Titanis most likely used its massive skull to pummel prey to death.[17][18]

Extinction

The extinction of T. walleri and other phorusrhacids throughout the Americas may have resulted from competition with large placental (canid, felid, and possibly ursid) carnivores that radiated in the same ancient terrestrial ecosystems during the Great American Interchange.[1]

From circumstantial evidence (i.e., bone fractures), it has been suggested that the species did not become extinct until 15,000 years ago,[19] but more precise dating by McFadden and colleagues refutes such a late date; all known Titanis fossils appear to be at least 2 million years old.[1][13]

References
  1. MacFadden, Bruce J.; Labs-Hochstein, Joann; Hulbert, Richard C.; Baskin, Jon A. (2007). "Revised age of the late Neogene terror bird (Titanis) in North America during the Great American Interchange". Geology. 35 (2): 123–126. Bibcode:2007Geo....35..123M. doi:10.1130/G23186A.1.
  2. Marsh, O. C. (1875). "On the Odontornithes, or birds with teeth". American Journal of Science. 10 (12): 403–408. Bibcode:1875AmJS...10..403M. doi:10.2475/ajs.s3-10.59.403. hdl:2027/hvd.32044107220493. JSTOR 2448315. S2CID 130972264.
  3. https://www.floridamemory.com/items/show/139737
  4. Dunbar, James (March–June 1983). "Florida Anthropologist Interview with Ben Waller". The Florida Anthropologist. 36 (Paleo–Indian): 31–39. Retrieved 22 June 2020.
  5. Garcia, Frank A. (1998). Discovering Fossils: How to Find and Identify Remains of the Prehistoric Past. Stackpole Books. pp. 86–87. ISBN 978-0811728003. Retrieved 22 June 2020.
  6. Degrange, Federico. "Society of Vertebrate Palaeontology".
  7. Richardson, Hazel (2003). Dinosaurs and Prehistoric Life. London: Dorling Kindersley Limited. p. 183. ISBN 0-7513-3734-X.
  8. Dyke, Gareth (15 February 2011). Living Dinosaurs: The Evolution of Modern Birds. John Wiley and Sons, Inc.
  9. Alvarenga, H. M. F.; Höfling, E. (2003). "Systematic revision of the Phorusrhacidae (Aves: Ralliformes)". Papéis Avulsos de Zoologia. 43 (4): 55–91. doi:10.1590/S0031-10492003000400001.
  10. Brodkorb, P. (1963). "A giant flightless bird from the Pleistocene of Florida" (PDF). Auk. 80 (2): 111–115. doi:10.2307/4082556. JSTOR 4082556.
  11. Chandler, R.M. (1994). "The wing of Titanis walleri (Aves: Phorusrhacidae) from the Late Blancan of Florida". Bulletin of the Florida Museum of Natural History, Biological Sciences. 36: 175–180.
  12. Gould, G.C. & Quitmyer, I.R. (2005). "Titanis walleri: bones of contention" (PDF). Bulletin of the Florida Museum of Natural History. 45: 201–229.
  13. McFadden, B.; Labs-Hochstein, J.; Hulbert Jr., R. C.; Baskin, J. A. (2006). "Refined age of the late Neogene terror bird (Titanis) from Florida and Texas using rare earth elements" (PDF). Journal of Vertebrate Paleontology. 26 (3): 92A (Supplement). doi:10.1080/02724634.2006.10010069. S2CID 220413406.
  14. Alroy, John, Ph.D. Synonymies and reidentifications of North American fossil mammals, 2002. John P. Hunter, Ohio State University, Mammalian Paleontology.
  15. Alvarenga, H.; Jones, W.; Rinderknecht, A. (May 2010). "The youngest record of phorusrhacid birds (Aves, Phorusrhacidae) from the late Pleistocene of Uruguay". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 256 (2): 229–234. doi:10.1127/0077-7749/2010/0052.
  16. Cracraft, Joel, A review of the Bathornithidae (Aves, Gruiformes), with remarks on the relationships of the suborder Cariamae. American Museum Novitates ; no. 2326
  17. Degrange, Federico J.; Tambussi, Claudia P.; Moreno, Karen; Witmer, Lawrence M.; Wroe, Stephen; Turvey, Samuel T. (18 August 2010). "Mechanical Analysis of Feeding Behavior in the Extinct "Terror Bird" Andalgalornis steulleti (Gruiformes: Phorusrhacidae)". PLOS ONE. 5 (8): e11856. Bibcode:2010PLoSO...511856D. doi:10.1371/journal.pone.0011856. PMC 2923598. PMID 20805872.
  18. Tambussi, Claudia P.; de Mendoza, Ricardo; Degrange, Federico J.; Picasso, Mariana B.; Evans, Alistair Robert (25 May 2012). "Flexibility along the Neck of the Neogene Terror Bird Andalgalornis steulleti (Aves Phorusrhacidae)". PLOS ONE. 7 (5): e37701. Bibcode:2012PLoSO...737701T. doi:10.1371/journal.pone.0037701. PMC 3360764. PMID 22662194.
  19. Baskin, J. A. (1995). "The giant flightless bird Titanis walleri (Aves: Phorusrhacidae) from the Pleistocene coastal plain of South Texas". Journal of Vertebrate Paleontology. 15 (4): 842–844. doi:10.1080/02724634.1995.10011266.
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