Colias palaeno

Colias palaeno, known by the common names moorland clouded yellow, palaeno sulphur, and pale Arctic clouded yellow, is a butterfly in the family Pieridae.

Moorland clouded yellow
C. p. palaeno, Sweden
C. p. europomene, female
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Lepidoptera
Family: Pieridae
Genus: Colias
Species:
C. palaeno
Binomial name
Colias palaeno
(Linnaeus, 1761)
Subspecies

Several, see text

Subspecies

Subspecies include:[1]

  • Colias palaeno palaeno – Sweden, Norway, Finland, and Estonia
  • Colias palaeno aias Fruhstorfer, 1903 – Japan the largest form of palaeno, the marginal band of the forewing being very broad and the underside rich green
  • Colias palaeno baffinensis Ebner & Ferris, [1978] – northern North America[2]
  • Colias palaeno chippewa W.H. Edwards, 1870 – northern North America,[3] may be a separate species (C. chippewa)[4]
  • Colias palaeno europome (Esper, 1778) – Belgium, Germany, Slovakia, Romania, and Ukraine
  • Colias palaeno europomene Ochsenheimer, 1808 – high altitude in the Alps. This form is somewhat larger than the nominate, the male lemon-yellow above, being below deeper yellow, as is also the female.
  • Colias palaeno orientalis Staudinger, 1892 – Kamchatka very similar to europomene, being dark greenish on the underside of the hindwing
  • Colias palaeno poktusani O. Bang-Haas, 1934 – North Korea
  • Colias palaeno sachalinensis Matsumura, 1919 – Sakhalin
  • Colias palaeno synonyma Bryk, 1923 – Sweden, Denmark[5]

Colias aias is treated as a full species by some authors.[6]

Distribution

Colias palaeno is a Holarctic species, widespread through Asia, Europe and North America. It is present in central and northern Europe from eastern France to the Baltic States and northern Sweden, Norway, Finland and to the eastern Europe, then in eastern Siberia, in the Chukchi Peninsula, in Japan and in northern areas of North America.[1][7][8]

Habitat

This species inhabits various types of moorland,[9] forest meadows, open coniferous forests[10] and open areas which may contain scrub. In the southern areas of its range it is a high alpine species reaching an elevation over 1,500 metres (4,900 ft) above sea level, but it can be found on upland bogs at an elevation of about 1,000 metres (3,300 ft).[11] Though the species is normally restricted to these habitats, occasionally it is found far from suitable breeding grounds.

Description
Colias palaeno

Colias palaeno has a wingspan of 46–50 millimetres (1.8–2.0 in) in males, of 50–52 millimetres (2.0–2.0 in) in females.[12] Upperside of male pale yellow with blackish brown distal margins, pale-centred dark middle spot to the forewing and light middle pot to the hindwing; fringes red. Underside of forewing pale yellow with white-centred dark middle spot, the costal and distal edges being red; hindwing yellow, strongly dusted with fuscous, the large middle spot being mother-of-pearl colour, and the fringes red. The female has a white ground colour above, the underside of the forewing being white proximally, yellow at apex, the hindwing being somewhat paler in the female than in the male.

The larva is sea-green, velvety, bearing minute black dots; a lateral stripe bright yellow edged with black beneath, below the same the white black-edged spiracles; underside and abdominal legs dull green, thoracic legs yellowish, head green. The Pupa is greenish yellow, the back being strongly convex.[13]

Biology

It is a univoltine species, flying from June to August.[9] As most Colias-species, Colias palaeno is an avid flyer. The larva is found on bog bilberry (Vaccinium uliginosum), on Vaccinium myrtillus and on Vaccinium caespitosum.[1]

  • Colias palaeno poktussani . Male, dorsal side
  • Female, dorsal side
  • Male, ventral side
  • Female, ventral side

References
  1. Funet at Markku Savela's Lepidoptera and Some Other Life Forms
  2. Colias palaeno baffinensis, Butterflies of America
  3. Colias palaeno chippewa, Butterflies of America
  4. Colias chippewa at Markku Savela's Lepidoptera and Some Other Life Forms
  5. Colias at Markku Savela's Lepidoptera and Some Other Life Forms
  6. Josef Grieshuber; Bob Worthy; Gerardo Lamas (2012), Münchner Entomologische Gesellschaft (ed.), The Genus Colias Fabricius, 1807: Jan Haugum'S Annotated Catalogue Of The Old World Colias (Lepidoptera, Pieridae) (in German), Pardubice: Tshikolovets Publications, ISBN 978-80-904900-2-4
  7. Delachaux et Niestlé, Tom Tolman, Richard Lewington, Guide des papillons d'Europe et d'Afrique du Nord (ISBN 978-2-603-01649-7)
  8. Fauna europaea
  9. Russian Insects
  10. Butterflies of the Asian part of Russia
  11. Matt Rowlings Eurobutterflies
  12. UK Butterflies
  13. Kimmo Silvonen Larvae of North-European Lepidoptera
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