2019 in paleoichthyology

This list of fossil fishes described in 2019 is a list of new taxa of jawless vertebrates, placoderms, acanthodians, fossil cartilaginous fishes, bony fishes, and other fishes of every kind that were described during the year 2019, as well as other significant discoveries and events related to paleoichthyology that occurred in 2019.

List of years in paleoichthyology
In paleontology
2016
2017
2018
2019
2020
2021
2022
In science
2016
2017
2018
2019
2020
2021
2022


New taxa

Jawless vertebrates

Name Novelty Status Authors Age Type locality Country Notes Images

Archegonaspis bashkirica[1]

Sp. nov

Valid

Märss

Silurian (Ludlow)

 Russia
( Bashkortostan)

A member of the family Cyathaspididae.

Cyathaspis alexanderi[1]

Sp. nov

Valid

Märss

Silurian

Tabuska Beds

 Russia
( Chelyabinsk Oblast)

A member of the family Cyathaspididae.

Jiaoyu[2]

Gen. et sp. nov

Valid

Liu et al.

Silurian

Kezirtag formation

 China

A member of Galeaspida of uncertain phylogenetic placement. The type species is J. imperfectus.

Parathelodus liaokuoensis[3]

Sp. nov

Valid

Cui et al.

Devonian (Lochkovian)

Xishancun Formation
Xitun Formation

 China

A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.

Parathelodus wangi[3]

Sp. nov

Valid

Cui et al.

Devonian (Lochkovian)

Xishancun Formation
Xitun Formation

 China

A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.

Parathelodus xitunensis[3]

Sp. nov

Valid

Cui et al.

Devonian (Lochkovian)

Xitun Formation

 China

A thelodont belonging to the family Coelolepidae. Announced in 2019; the final version of the article naming it was published in 2020.

Tethymyxine[4]

Gen. et sp. nov

Valid

Miyashita et al.

Late Cretaceous (Cenomanian)

Hjoula Lagerstätte

 Lebanon

A hagfish. The type species is T. tapirostrum.

Tolypelepis bedovensis[1]

Sp. nov

Valid

Märss

Late Silurian

Krasnaya Bukhta Formation

 Russia
( Krasnoyarsk Krai)

A member of the family Cyathaspididae.

Xiyuaspis[2]

Gen. et comb. nov

Valid

Liu et al.

Silurian

Tataertag Formation

 China

A member of Galeaspida belonging to the group Hanyangaspidida and to the family Xiushuiaspidae. The type species is "Nanjiangaspis" zhangi Lu et al. (2007).

Placoderms

Name Novelty Status Authors Age Type locality Country Notes Images

Asterolepis alticristata[5]

Sp. nov

Valid

Downs et al.

Devonian (Frasnian)

Fram Formation

 Canada
( Nunavut)

An antiarch in Asterolepis

Colombialepis[6]

Gen. et sp. nov

Valid

Olive et al.

Late Devonian

Cuche Formation

 Colombia

A member of Antiarchi, superfamily Asterolepidoidei. Genus includes new species C. villarroeli.

Colombiaspis[6]

Gen. et sp. nov

Valid

Olive et al.

Late Devonian

Cuche Formation

 Colombia

A member of Arthrodira, family Groenlandaspidae. Genus includes new species C. rinconensis.

Meeksiella[7]

Gen. et sp. nov

Valid

Trinajstic et al.

Devonian (Frasnian)

Snetnya Gora Beds

 Estonia
 Latvia
 Russia
( Pskov Oblast)

A member of Ptyctodontida. The type species is "Chelyophorus" pskovensis Obruchev (1947).

Acanthodians

Name Novelty Status Authors Age Type locality Country Notes Images

Cheiracanthus peachi[8]

Sp. nov

Valid

Den Blaauwen, Newman & Burrow

Devonian (Givetian)

Mey Flagstone Formation
Rousay Flagstone Formation

 United Kingdom

Serradentus[9]

Gen. et sp. nov

Newman, Burrow & Den Blaauwen

Devonian (Givetian)

Tordalen Formation

 Germany
 Norway

A member of Ischnacanthiformes. The type species is S. armstrongi.

Cartilaginous fishes

Name Novelty Status Authors Age Type locality Country Notes Images

Antiquaobatis[10]

Gen. et sp. nov

Valid

Stumpf & Kriwet

Early Jurassic (Pliensbachian)

 Germany

A member of Rajiformes of uncertain phylogenetic placement. The type species is A. grimmenensis.

Asflapristis[11]

Gen. et sp. nov

Valid

Villalobos-Segura et al.

Late Cretaceous (Turonian)

Akrabou Formation

 Morocco

A member of Sclerorhynchoidei belonging to the family Ptychotrygonidae. Genus includes new species A. cristadentis.

Cantioscyllium clementsi[12]

Sp. nov

Valid

Case et al.

Late Cretaceous (Campanian)

Bladen Formation

 United States
( North Carolina)

A member of the family Ginglymostomatidae.

Carcharhinus mancinae[13]

Sp. nov

Valid

Ebersole, Cicimurri & Stringer

Eocene (Bartonian)

Gosport Sand
Lisbon Formation

 United States
( Alabama)

A species of Carcharhinus.

Carcharhinus underwoodi[14]

Sp. nov

Valid

Samonds et al.

Eocene

Mahajanga Basin

 Madagascar

A species of Carcharhinus.

Cretalamna feldmanni[15]

Sp. nov

Valid

Hoganson, Erickson & Holland

Late Cretaceous (Maastrichtian)

Fox Hills Formation

 United States
( North Dakota)

Cretodus houghtonorum[16]

Sp. nov

Valid

Shimada & Everhart

Late Cretaceous (Turonian)

Carlile Shale

 United States
( Kansas)

A mackerel shark belonging to the family Pseudoscapanorhynchidae.

Dasyatis northdakotaensis[15]

Sp. nov

Valid

Hoganson, Erickson & Holland

Late Cretaceous (Maastrichtian)

Fox Hills Formation

 United States
( North Dakota)

A species of Dasyatis.

Diprosopovenator[17]

Gen. et sp. nov

Valid

Stumpf, Scheer & Kriwet

Late Cretaceous (Cenomanian)

Hesseltal Formation

 Germany

A member of Carcharhiniformes belonging to the new family Pseudoscyliorhinidae. The type species is D. hilperti.

Galagadon[18]

Gen. et sp. nov

Valid

Gates, Gorscak & Makovicky

Late Cretaceous (Maastrichtian)

Hell Creek Formation

 United States
( South Dakota)

A carpet shark. The type species is G. nordquistae.

Glencartius[19]

Gen. et comb. nov

Valid

Ginter & Skompski

Carboniferous (Viséan)

 Poland
 United Kingdom

A member of Ctenacanthiformes of uncertain phylogenetic placement. The type species is "Ctenacanthus" costellatus Traquair (1884).

Gydoselache[20]

Gen. et sp. nov

Valid

Maisey et al.

Devonian (Emsian)

Gydo Formation

 South Africa

A cartilaginous fish of uncertain phylogenetic placement, related to Pucapampella and assigned to the new family Pucapampellidae. The type species is G. oosthuizeni.

Hypolophites beckeri[21]

Sp. nov

Valid

Maisch

Paleocene

Midway Group

 United States
( Arkansas)

A member of Myliobatiformes.

Isogomphodon aikenensis[22]

Sp. nov

Valid

Cicimurri & Knight

Eocene (Priabonian)

Dry Branch Formation

 United States
( South Carolina)

A relative of the daggernose shark.

Kimmerobatis[23]

Gen. et sp. nov

Valid

Underwood & Claeson

Late Jurassic (Kimmeridgian)

Kimmeridge Clay

 United Kingdom

A member of Batoidea related to Spathobatis. Genus includes new species K. etchesi.

Lessiniabatis[24]

Gen. et sp. nov

Valid

Marramà et al.

Eocene (Ypresian)

Monte Bolca Konservat-Lagerstätte

 Italy

A member of Myliobatiformes belonging to the superfamily Dasyatoidea. The type species is L. aenigmatica.

Myledaphus araucanus[25]

Sp. nov

Valid

Otero

Late Cretaceous (Maastrichtian)

Arauco Basin

 Chile

"Myliobatis" foxhillsensis[15]

Sp. nov

Valid

Hoganson, Erickson & Holland

Late Cretaceous (Maastrichtian)

Fox Hills Formation

 United States
( North Dakota)

An eagle ray.

Odontorhytis bahariensis[26]

Sp. nov

Valid

Salame & Asan

Eocene (probably Lutetian)

 Egypt

A member of Neoselachii of uncertain phylogenetic placement.

Ottangodus[27]

Gen. et sp. nov

Valid

Popov, Delsate & Felten

Middle Jurassic (Bajocian)

Luxembourg-French border

A chimaera belonging to the family Callorhinchidae. Genus includes new species O. lotharingiae.

Pastinachus kebarensis[28]

Sp. nov

Valid

Adnet et al.

Eocene (late Bartonian)

 Tunisia

A cowtail stingray of Pastinachus. Announced in 2018; the final version of the article naming it was published in 2019.

Pristiophorus humboldti[29]

Sp. nov

Valid

Villafaña et al.

Early Miocene

 Chile

A species of Pristiophorus.

Protolamna ricaurtensis[30]

Sp. nov

Valid

Carrillo-Briceño, Parra & Luque

Early Cretaceous (late Barremian–early Aptian)

Paja Formation

 Colombia

A mackerel shark belonging to the family Pseudoscapanorhynchidae.

Pseudabdounia[13]

Gen. et comb. nov

Valid

Ebersole, Cicimurri & Stringer

Eocene

Gosport Sand
Lisbon Formation
Tallahatta Formation

 Belgium
 United States
( Alabama)

A requiem shark. The type species is "Galeocerdo" recticonus Winkler (1874); genus also includes "Galeorhinus recticonus" claibornensis White (1956) (raised to the rank of a separate species Pseudabdounia claibornensis).

Ptychotrygon rostrispatula[31]

Sp. nov

Valid

Villalobos‐Segura, Underwood & Ward

Late Cretaceous (Turonian)

 Morocco

Scymnodalatias kazenobon[32]

Sp. nov

Valid

Nishimatsu & Ujihara

Middle Miocene

Yatsuo Group

 Japan

A species of Scymnodalatias.

Tethytrygon[33]

Gen. et comb. nov

Valid

Marramà et al.

Eocene (Ypresian)

Monte Bolca

 Italy

A whiptail stingray belonging to the subfamily Neotrygoninae; a new genus for "Raja" muricata Volta (1796).

Tlalocbatos[34]

Gen. et sp. nov

Valid

Brito, Villalobos-Segura & Alvarado-Ortega

Early Cretaceous

Tlayúa Formation

 Mexico

A member of Batoidea related to the banjo rays. Genus includes new species T. applegatei.

Ray-finned fishes

Name Novelty Status Authors Age Type locality Country Notes Images

Acronuroides[35]

Gen. et sp. nov.

Valid

Bannikov, Carnevale & Tyler

Eocene (late Ypresian)

Monte Bolca locality

 Italy

A member of Percomorphacea of uncertain phylogenetic placement. The type species is A. eocaenicus.

Acropoma aurora[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan to Bortonian

 New Zealand

A species of Acropoma.

Ambassis simesi[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

A species of Ambassis.

Ampheristus pentlandensis[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

A member of the family Ophidiidae.

Antigonia artata[36]

Sp. nov

Valid

Schwarzhans

Waipawan-Heretaungan

 New Zealand

A species of Antigonia.

Antimyctophum[36]

Gen. et comb. nov

Valid

Schwarzhans

Clifdenian to Tongaporutuan

 New Zealand

A lanternfish. The type species is "Scopelus" konganaruensis Frost (1933).

Archoplites langrellorum[37]

Sp. nov

Valid

Van Tassell & Smith

Pliocene (Blancan)

 United States
( Oregon)

A relative of the Sacramento perch.

Ardoreosomus[38]

Gen. et sp. nov

Valid

Romano et al.

Early Triassic (Induan)

Candelaria Formation

 United States
( Nevada)

A member of the family Ptycholepidae. Genus includes new species A. occidentalis.

Argyripnus oamaruensis[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Argyripnus.

Argyripnus rapahoensis[36]

Sp. nov

Valid

Schwarzhans

Runangan

 New Zealand

A species of Argyripnus.

Arnoglossus lautus[36]

Sp. nov

Valid

Schwarzhans

Duntroonian and Waitakian

 New Zealand

A scaldfish.

Arnoglossus purus[36]

Sp. nov

Valid

Schwarzhans

Lillburnian and Waiauan

 New Zealand

A scaldfish.

Artediellichthys candelabrum[39]

Sp. nov

Valid

Nazarkin

Miocene

Agnevo Formation

 Russia
( Sakhalin Oblast)

A relative of the blackfin hookear sculpin.

Artediellus simplex[39]

Sp. nov

Valid

Nazarkin

Miocene

Agnevo Formation

 Russia
( Sakhalin Oblast)

A species of Artediellus.

Aseraggodes hudsoni[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Aseraggodes.

Auchenoceros simplex[36]

Sp. nov

Valid

Schwarzhans

Duntroonian

 New Zealand

A relative of the ahuru.

Aulichthys miocaenicus[40]

Sp. nov

Valid

Nazarkin

Miocene (SerravallianTortonian)

Agnevo Formation

 Russia
( Sakhalin Oblast)

An extinct tubenose.

Ausonasynodus[41]

Gen. et sp. nov

Valid

Carnevale et al.

Late Eocene

 Spain

A lizardfish. Genus includes new species A. almerai.

Awamoa[36]

Gen. et comb. et sp. nov

Valid

Schwarzhans

Otaian to Opoitian

 New Zealand

A southern sandfish. The type species is "Leptoscopus" progressus Schwarzhans (1980); genus also includes "Leptoscopus" iocosus Schwarzhans (1980) and a new species A. kaawa.

Balistes vegai[42]

Sp. nov

Valid

Viñola Lopez, Carr & Lorenzo

Miocene

 Cuba

A species of Balistes.

Barathronus nielseni[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Barathronus.

Barbalepis[43]

Gen. et comb. nov

Valid

Olive, Taverne & López-Arbarello

Early Cretaceous (BarremianAptian)

 Belgium

A member the family Coccolepididae; a new genus for "Coccolepis" macroptera Traquair.

Bathycongrus waihaoensis[36]

Sp. nov

Valid

Schwarzhans

Kaiatan

 New Zealand

A species of Bathycongrus.

Bathygadus waiohaensis[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Bathygadus.

Bembrops aequiformis[36]

Sp. nov

Valid

Schwarzhans

Duntroonian and Waitakian

 New Zealand

A species of Bembrops.

Bembrops grenfelli[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Bembrops.

Benthosema minutum[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Benthosema.

Bidenichthys struthersi[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Bidenichthys.

Blachea semeniformis[36]

Sp. nov

Valid

Schwarzhans

Waitakian to Altonian

 New Zealand

A species of Blachea.

Bonapartia altidorsalis[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A member of the family Gonostomatidae.

Brosmophyciops tongarewae[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan-Heretaungan

 New Zealand

A species of Brosmophyciops.

Burguklia minichorum[44]

Sp. nov

Valid

Bakaev & Kogan

Permian (WordianCapitanian)

 Russia

An early ray-finned fish. Announced in 2019; the final version of the article naming it was published in 2020.

Cabindachanos[45]

Gen. et sp. nov

Valid

Taverne et al.

Paleocene (Danian or early Selandian)

 Angola

A member of the family Chanidae. The type species is C. dartevellei.

Callionymus hakatarameaensis[36]

Sp. nov

Valid

Schwarzhans

Waitakian

 New Zealand

A species of Callionymus.

Callionymus triquetrus[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Callionymus.

Candelarialepis[38]

Gen. et sp. nov

Valid

Romano et al.

Early Triassic (Induan)

Candelaria Formation

 United States
( Nevada)

A stem-neopterygian belonging to the family Parasemionotidae. Genus includes new species C. argentus.

Cavinichthys[46]

Gen. et sp. nov

Valid

Taverne & Capasso

Early Cretaceous (Albian)

Limestones of Pietraroja

 Italy

A member of Crossognathiformes belonging to the family Pachyrhizodontidae. The type species is C. pachylepis.

Cephalopholis? aotearoa[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan to Bortonian

 New Zealand

Possibly a species of Cephalopholis.

Cepola ambifaria[36]

Sp. nov

Valid

Schwarzhans

Otaian to Clifdenian

 New Zealand

A species of Cepola.

Ceratoscopelus richardsoni[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Ceratoscopelus.

Champsodon timaruensis[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Champsodon.

Chaunax bucculentus[36]

Sp. nov

Valid

Schwarzhans

Waitakian

 New Zealand

A species of Chaunax.

Chaychanus[47]

Gen. et sp. nov

Valid

Cantalice Severiano, Alvarado Ortega & Bellwood

Paleocene

 Mexico

A member of the family Pomacentridae. Genus includes new species C. gonzalezorum. Announced in 2019; the final version of the article naming it was published in 2020.

Chrionema salebrosa[36]

Sp. nov

Valid

Schwarzhans

Duntroonian to Altonian

 New Zealand

A species of Chrionema.

Coelorinchus divulgatus[36]

Sp. nov

Valid

Schwarzhans

Waitakian to Lillburnian

 New Zealand

A species of Coelorinchus.

Coelorinchus pakaurangiensis[36]

Sp. nov

Valid

Schwarzhans

Duntroonian to Otaian

 New Zealand

A species of Coelorinchus.

Coelorinchus preaustralis[36]

Sp. nov

Valid

Schwarzhans

Duntroonian to Altonian

 New Zealand

A species of Coelorinchus.

Colobodus wushaensis[48]

Sp. nov

Valid

Li et al.

Late Triassic (Carnian)

Falang Formation

 China

A member of Perleidiformes.

Conger davidsmithi[36]

Sp. nov

Valid

Schwarzhans

Waipawan to Bortonian

 New Zealand

A species of Conger.

Conger tokoroa[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Conger.

Congopycnodus[49]

Gen. et sp. nov

Valid

Taverne

Middle Jurassic

Stanleyville Formation

 Democratic Republic of the Congo

A member of Pycnodontiformes belonging to the superfamily Coccodontoidea. The type species is C. cornutus.

Costapycnodus[50]

Gen. et comb. nov

Valid

Taverne, Capasso & Del Re

Early Cretaceous (late Hauterivian-early Barremian)

 Italy

A member of the family Pycnodontidae. The type species is "Coelodus" costae Heckel (1856).

Danaphos glomerosus[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Danaphos.

Diaphus audax[36]

Sp. nov

Valid

Schwarzhans

Waitakian to Altonian

 New Zealand

A species of Diaphus.

Diaphus caurus[51]

Sp. nov

Valid

Schwarzhans & Ohe

Pleistocene (Calabrian)

Hijikata Formation

 Japan

A species of Diaphus.

Diaphus endoi[51]

Sp. nov

Valid

Schwarzhans & Ohe

Pliocene (Piacenzian)

Shinzato Formation

 Japan

A species of Diaphus.

Diaphus exilis[36]

Sp. nov

Valid

Schwarzhans

Otaian to Lillburnian

 New Zealand

A species of Diaphus.

Diaphus grebneffi[51]

Sp. nov

Valid

Schwarzhans & Ohe

Probably Piacenzian

Nakosi Formation

 Fiji

A species of Diaphus.

Diaphus huatau[36]

Sp. nov

Valid

Schwarzhans

Clifdenian to Lillburnian

 New Zealand

A species of Diaphus.

Diaphus kaiparaensis[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Diaphus.

Diaphus kakegawaensis[51]

Sp. nov

Valid

Schwarzhans & Ohe

Pleistocene (Gelasian)

Dainichi Formation

 Japan

A species of Diaphus.

Diaphus manneringi[36]

Sp. nov

Valid

Schwarzhans

Clifdenian to Waiauan

 New Zealand

A species of Diaphus.

Diaphus mirus[36]

Sp. nov

Valid

Schwarzhans

Tongaporutuan

 New Zealand

A species of Diaphus.

Diaphus nafpaktitisi[51]

Sp. nov

Valid

Schwarzhans & Ohe

Pliocene (Piacenzian)

Nobori Formation

 Japan

A species of Diaphus.

Diaphus noboriensis[51]

Sp. nov

Valid

Schwarzhans & Ohe

Pliocene (Piacenzian) and Pleistocene (Gelasian)

Dainichi Formation
Nobori Formation
Shinzato Formation

 Japan

A species of Diaphus.

Diaphus tenax[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Diaphus.

Dibranchus kakahoensis[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

A species of Dibranchus.

Diogenichthys rangiauriensis[36]

Sp. nov

Valid

Schwarzhans

Mangapanian

 New Zealand

A species of Diogenichthys.

Dipulus certus[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan-Heretaungan

 New Zealand

A species of Dipulus.

Dolichopteryx iustus[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Dolichopteryx.

Echiodon teres[36]

Sp. nov

Valid

Schwarzhans

Otaian to Clifdenian

 New Zealand

A species of Echiodon.

Electrona subasperoides[36]

Sp. nov

Valid

Schwarzhans

Waitakian to Altonian

 New Zealand

A species of Electrona.

Ellimma longipectoralis[52]

Sp. nov

Valid

Polck et al.

Early Cretaceous (Aptian)

Barra Velha Formation

 Brazil

A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Paraclupeidae. Announced in 2019; the final version of the article naming it was published in 2020.

Emmelichthys tennysoni[36]

Sp. nov

Valid

Schwarzhans

Waitakian

 New Zealand

A species of Emmelichthys.

Encheliophis strigosus[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Encheliophis.

Eomyctophum broncus[36]

Sp. nov

Valid

Schwarzhans

Waipawan to Porangan

 New Zealand

A lanternfish.

Eomyctophum porokawa[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

A lanternfish.

Eosemionotus diskosomus[53]

Sp. nov

Valid

López-Arbarello et al.

Middle Triassic (Ladinian)

Meride Limestone

 Italy
  Switzerland

A member of Semionotiformes.

Eosemionotus minutus[53]

Sp. nov

Valid

López-Arbarello et al.

Middle Triassic (Ladinian)

Meride Limestone

  Switzerland

A member of Semionotiformes.

Eosemionotus sceltrichensis[53]

Sp. nov

Valid

López-Arbarello et al.

Middle Triassic (Ladinian)

Meride Limestone

  Switzerland

A member of Semionotiformes.

Epaelops[54]

Gen. et sp. nov

Valid

Alves, Alvarado-Ortega & Brito

Early Cretaceous (Albian)

Tlayúa Formation

 Mexico

A member of Elopiformes. Genus includes new species E. martinezi. Announced in 2019; the final version of the article naming it was published in 2020.

Epigonus aquilonius[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Epigonus.

Epigonus opoitiensis[36]

Sp. nov

Valid

Schwarzhans

Opoitian

 New Zealand

A species of Epigonus.

Etrumeus carnatus[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Etrumeus.

Evynnis abax[36]

Sp. nov

Valid

Schwarzhans

Duntroonian to Altonian

 New Zealand

A member of the family Sparidae.

Flagellipinna[55]

Gen. et sp. nov

Valid

Cawley & Kriwet

Late Cretaceous (Cenomanian)

Sannine Formation

 Lebanon

A member of Pycnodontiformes belonging to the family Pycnodontidae. The type species is F. rhomboides.

Forsterygion jawadi[36]

Sp. nov

Valid

Schwarzhans

Nukumaruan

 New Zealand

A species of Forsterygion.

Gadiculus adversus[36]

Sp. nov

Valid

Schwarzhans

Waitakian to Lillburnian

 New Zealand

A species of Gadiculus.

Gadiculus pereawa[36]

Sp. nov

Valid

Schwarzhans

Tongaporutuan

 New Zealand

A species of Gadiculus.

Galeichthys ohei[36]

Sp. nov

Valid

Schwarzhans

Opoitian to Nukumaruan

 New Zealand

A species of Galeichthys.

Gilmourella[56]

Gen. et sp. nov.

Valid

Carnevale & Bannikov

Eocene (late Ypresian)

Monte Bolca locality

 Italy

A member of Callionymoidei. The type species is G. minuta.

Gnathophis araroa[36]

Sp. nov

Valid

Schwarzhans

Kapitean

 New Zealand

A species of Gnathophis.

Gonostoma retunsum[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Gonostoma.

Grahamichthys frigophila[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A member of the family Thalasseleotrididae.

Guus[57]

Gen. et comb. nov.

Valid

Bannikov

Eocene (late Ypresian)

Monte Bolca locality

 Italy

A member of the family Tortonesidae. The type species is "Gobius" microcephalus Agassiz (1839).

Gyrodus huiliches[58]

Sp. nov

Valid

Gouiric-Cavalli, Remírez & Kriwet

Early Cretaceous (Valanginian–early Hauterivian)

Agrio Formation

 Argentina

Harpagifer? morgansi[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

Possibly a species of Harpagifer.

Hemerocoetes pukunati[36]

Sp. nov

Valid

Schwarzhans

Duntroonian to Altonian

 New Zealand

A species of Hemerocoetes.

Hemerocoetes whiroki[36]

Sp. nov

Valid

Schwarzhans

Duntroonian to Altonian

 New Zealand

A species of Hemerocoetes.

Hiascoactinus[59]

Gen. et sp. nov

Valid

Kim et al.

Late Triassic

Amisan Formation

 South Korea

A member of Redfieldiiformes. Genus includes new species H. boryeongensis. Announced in 2019; the final version of the article naming it was published in 2020.

Hintonia robertsi[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A lanternfish.

Howella monodens[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Howella.

Hygophum acutiventris[36]

Sp. nov

Valid

Schwarzhans

Kapitean

 New Zealand

A species of Hygophum.

Ichthyscopus pukeuriensis[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Ichthyscopus.

Klincigobius[60]

Gen. et comb. et sp. nov

Valid

Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al.

Early Miocene

 Serbia

A member of the family Gobiidae. The type species is "Gobius" serbiensis Gaudant (1998); genus also includes new species K. andjelkovicae.

Kradimus[61]

Gen. et sp. nov

Valid

Veysey, Brito & Martill

Late Cretaceous (Turonian)

Akrabou Formation

 Morocco

A member of Crossognathiformes. Genus includes new species K. asflaensis. Announced in 2019; the final version of the article naming it was published in 2020.

Lactarius primigenius[36]

Sp. nov

Valid

Schwarzhans

Bortonian to Kaiatan

 New Zealand

A relative of the false trevally.

Lactarius pusillus[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A relative of the false trevally.

Laeops undulatus[36]

Sp. nov

Valid

Schwarzhans

Lillburnian

 New Zealand

A species of Laeops.

Lampanyctodes transitivus[36]

Sp. nov

Valid

Schwarzhans

Kapitean

 New Zealand

A lanternfish.

Lampanyctus popoto[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Lampanyctus.

Lampanyctus profestus[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Lampanyctus.

Lampanyctus waiohaensis[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Lampanyctus.

Lashanichthys[62]

Gen. et comb. nov

Valid

Xu et al.

Middle Triassic (Anisian)

 China

A member of Ginglymodi assigned to the group Kyphosichthyiformes and to the new family Lashanichthyidae. The type species is "Sangiorgioichthys" sui López-Arbarello et al. (2011); genus also includes "Sangiorgioichthys" yangjuanensis Chen et al. (2014).

Latellopsis[63]

Gen. et comb. nov.

Valid

Bannikov & Zorzin

Eocene (late Ypresian)

Monte Bolca locality

 Italy

A member of Perciformes of uncertain phylogenetic placement. The type species is "Psettopsis" latellai Bannikov (2005).

Lepidorhynchus frosti[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A member of the family Macrouridae.

Leptoscopus atavus[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A southern sandfish.

Lethrinus? crassiornatus[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan-Heretaungan

 New Zealand

Possibly a species of Lethrinus.

Lindoeichthys[64]

Gen. et sp. nov.

Valid

Murray et al.

Late Cretaceous (Maastrichtian)

Scollard Formation

 Canada
( Alberta)

A member of Percopsiformes. Genus includes new species L. albertensis. Announced in 2019; the final version of the article naming it was published in 2020.

Liparis? hampdenensis[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

Possibly a species of Liparis.

Loancorhynchus[65]

Gen. et sp. nov

Valid

Otero

Middle Eocene

Millongue Formation

 Chile

A relative of the swordfish. The type species is L. catrillancai.

Lophiodes hoi[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Lophiodes.

Lophius alveolus[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Lophius.

Lophius onokensis[36]

Sp. nov

Valid

Schwarzhans

Mangapanian to Nukumaruan

 New Zealand

A species of Lophius.

Lotella matata[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Lotella.

Lotella pfeili[36]

Sp. nov

Valid

Schwarzhans

Altonian and Clifdenian

 New Zealand

A species of Lotella.

Lotella spicata[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan-Heretaungan

 New Zealand

A species of Lotella.

Macrurulus depressus[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

A member of the family Merlucciidae.

Macrurulus fordycei[36]

Sp. nov

Valid

Schwarzhans

Waipawan-Mangaorapan

 New Zealand

A member of the family Merlucciidae.

Maoricottus[36]

Gen. et comb. et 2 sp. nov

Valid

Schwarzhans

Altonian to Lillburnian

 New Zealand

A member of the family Cottidae. The type species is "Cottidarum" impolitus Schwarzhans (1980); genus also includes new species M. calidophilus and M. ovatus.

Melamphaes leeae[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Melamphaes.

Moradebrichthys[66]

Gen. et sp. nov

Valid

Cartanyà et al.

Middle Triassic (Ladinian)

 Spain

A member of the family Perleididae. Genus includes new species M. vilasecae.

Muraenanguilla[36]

Gen. et comb. et 2 sp. nov

Valid

Schwarzhans

Paleocene and Eocene

 Belgium
 New Zealand

A member of Anguilloidei of uncertain phylogenetic placement. The type species is Otolithus (Trachini)" thevenini Priem (1906); genus also includes new species M. balegemensis and M. lacinata.

Myctophum bortonensis[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

A species of Myctophum.

Myctophum tamumuensis[36]

Sp. nov

Valid

Schwarzhans

Clifdenian to Waiauan

 New Zealand

A species of Myctophum.

Myctophum tenellum[36]

Sp. nov

Valid

Schwarzhans

Tongaporutuan

 New Zealand

A species of Myctophum.

Mystocheilus[37]

Gen. et sp. nov

Valid

Van Tassell & Smith

Pliocene (Blancan)

 United States
( Oregon)

A member of the family Cyprinidae. The type species is M. fresti.

Nemadactylus trulliformis[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Nemadactylus.

Nemadactylus utoka[36]

Sp. nov

Valid

Schwarzhans

Waitakian

Otekaike Limestone

 New Zealand

A species of Nemadactylus.

Neobythites turpidus[36]

Sp. nov

Valid

Schwarzhans

Runangan

 New Zealand

A species of Neobythites.

Nezumia morgansi[36]

Sp. nov

Valid

Schwarzhans

Bortonian and Kaiatan

 New Zealand

A species of Nezumia.

Notoconger[36]

Gen. et 2 sp. nov

Valid

Schwarzhans

Runangan to Altonian

 New Zealand

A member of the family Congridae. The type species is N. hesperis; genus also includes N. devexus.

Notoscopelus effertus[36]

Sp. nov

Valid

Schwarzhans

Kapitean

 New Zealand

A species of Notoscopelus.

Notoscopelus praejaponicus[51]

Sp. nov

Valid

Schwarzhans & Ohe

Pleistocene (Calabrian)

Chinen Formation
Hijikata Formation

 Japan

A species of Notoscopelus.

Opistognathus wharekuriensis[36]

Sp. nov

Valid

Schwarzhans

Duntroonian

 New Zealand

A species of Opistognathus.

Oreochromimos[67]

Gen. et sp. nov

Penk et al.

Miocene

Ngorora Formation

 Kenya

A cichlid belonging to the tribe Oreochromini. The type species is O. kabchorensis.

Padangia[68]

Gen. et comb. nov

Valid

Murray

Probably Eocene

 Indonesia

A member of the family Cyprinidae. Genus includes "Sardinioides" amblyostoma von der Marck (1876).

Panturichthys grenfelli[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Panturichthys.

Paralabrus[69]

Gen. et sp. nov.

Valid

Bannikov & Zorzin

Eocene (late Ypresian)

Monte Bolca locality

 Italy

Possibly a wrasse. The type species is P. rossiae.

Parapercis bispicatus[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Parapercis.

Parapercis depressidorsalis[36]

Sp. nov

Valid

Schwarzhans

Waitakian to Altonian

 New Zealand

A species of Parapercis.

Parapercis fatoides[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Parapercis.

Parapercis pareoraensis[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Parapercis.

Parapercis richardsoni[36]

Sp. nov

Valid

Schwarzhans

Waipawan to Porangan

 New Zealand

A species of Parapercis.

Parapercis waiwaia[36]

Sp. nov

Valid

Schwarzhans

Opoitian

 New Zealand

A species of Parapercis.

Parascombrops giganteus[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan

 New Zealand

A member of the family Acropomatidae.

Parascombrops schwarzhansi[70]

Sp. nov

Valid

Van Hinsbergh & Helwerda

Late Pliocene to early Pleistocene

 Philippines

A member of the family Acropomatidae.

Paraulopus rallus[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

A species of Paraulopus.

Pelargorhynchus grandis[71]

Sp. nov

Wallaard et al.

Late Cretaceous (Maastrichtian)

Maastricht Formation

 Netherlands

A member of Aulopiformes belonging to the family Dercetidae.

Pempheris hurupiensis[36]

Sp. nov

Valid

Schwarzhans

Tongaporutuan

Hurupi Formation

 New Zealand

A species of Pempheris.

Phractocephalus yaguaron[72]

Sp. nov

Valid

Bogan & Agnolín

Late Miocene

 Argentina

A relative of the redtail catfish.

Physiculus beui[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Physiculus.

Platycephalus? iaiunus[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan to Bortonian

 New Zealand

Possibly a species of Platycephalus.

Platycephalus? riremoana[36]

Sp. nov

Valid

Schwarzhans

Runangan

 New Zealand

Possibly a species of Platycephalus.

Platysiagum sinensis[73]

Sp. nov

Valid

Wen et al.

Middle Triassic (Anisian)

Guanling Formation

 China

Protomyctophum ahunga[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Protomyctophum.

Pseudaequalobythites[36]

Gen. et sp. et comb. nov

Valid

Schwarzhans

Paleocene and Eocene

 Belgium
 France
 Germany
 New Zealand
 United Kingdom

A member of the family Ophidiidae. The type species is P. biplex; genus also includes "Otolithus" hilgendorfi Koken (1891).

Pseudanthias multicrenatus[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan to Bortonian

 New Zealand

A species of Pseudanthias.

Pseudocaranx? pertenuis[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

Possibly a species of Pseudocaranx.

Pseudophycis muringa[36]

Sp. nov

Valid

Schwarzhans

Lillburnian

 New Zealand

A species of Pseudophycis.

Pterygotrigla stewarti[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Pterygotrigla.

Pteronisculus nevadanus[38]

Sp. nov

Valid

Romano et al.

Early Triassic (Induan)

Candelaria Formation

 United States
( Nevada)

Possibly a member of the family Turseoidae.

Pycnodus multicuspidatus[74]

Sp. nov

Valid

Vullo et al.

Paleocene (Thanetian)

 Morocco

Quasinectes[63]

Gen. et sp. nov.

Valid

Bannikov & Zorzin

Eocene (late Ypresian)

Monte Bolca locality

 Italy

A member of Perciformes of uncertain phylogenetic placement. The type species is Q. durello.

Rebekkachromis[75]

Gen. et 2 sp. nov

Valid

Kevrekidis, Valtl & Reichenbacher in Kevrekidis et al.

Miocene

Ngorora Formation

 Kenya

A cichlid belonging to the subfamily Pseudocrenilabrinae and the tribe Oreochromini.[76] The type species is R. ngororus; genus also includes R. kiptalami.

Rhamphogobius[60]

Gen. et sp. et comb. nov

Valid

Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al.

Early Miocene

 Croatia
 Germany
 Serbia
  Switzerland

A member of the family Gobiidae. The type species is R. varidens; genus also includes "Gobius" doppleri Reichenbacher (1993), "Gobius" gregori Reichenbacher (1993) and "Gobius" helvetiae Salis (1967).

Rhynchoconger otaianus[36]

Sp. nov

Valid

Schwarzhans

Waipawan-Mangaorapan

 New Zealand

A species of Rhynchoconger.

Saccogaster parengarenga[36]

Sp. nov

Valid

Schwarzhans

Otaian to Clifdenian

 New Zealand

A species of Saccogaster.

Sardinella claviformis[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Sardinella.

Sardinella lintriculus[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Sardinella.

Schernfeldfuro[77]

Gen. et sp. nov

Valid

Ebert

Late Jurassic

 Germany

A member of Halecomorphi. Genus includes new species S. uweelleri.

Scleropages sanshuiensis[78]

Sp. nov

Valid

Zhang

Early Eocene

Huachong Formation

 China

A species of Scleropages. Announced in 2019; the final version of the article naming it was published in 2020.

Scopelarchoides neamticus[79]

Sp. nov

Valid

Grădianu et al.

Oligocene

Lower Dysodilic Shales Formation

 Romania

A pearleye, a species of Scopelarchoides.

Scopelosaurus? brevicauda[36]

Sp. nov

Valid

Schwarzhans

Kaiatan

 New Zealand

Possibly species of Scopelosaurus.

Sillaginodes albisaxosus[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A member of the family Sillaginidae.

Sillago maxwelli[36]

Sp. nov

Valid

Schwarzhans

Bortonian

 New Zealand

A species of Sillago.

Sprattus arewhana[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Sprattus.

Sundabarbus[68]

Gen. et comb. nov

Valid

Murray

Probably Eocene

Sangkarewang Formation

 Indonesia

A member of the family Cyprinidae. Genus includes "Barbus" megacephalus Günther (1876).

Symbolophorus moriguchii[51]

Sp. nov

Valid

Schwarzhans & Ohe

Pliocene (Piacenzian) and Pleistocene (Gelasian)

Dainichi Formation
Nobori Formation
Shinzato Formation

 Japan

A species of Symbolophorus.

Symbolophorus opononiensis[36]

Sp. nov

Valid

Schwarzhans

Waitakian and Otaian

 New Zealand

A species of Symbolophorus.

Symbolophorus tongaporutuensis[36]

Sp. nov

Valid

Schwarzhans

Tongaporutuan

 New Zealand

A species of Symbolophorus.

Symphysanodon inamata[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan to Bortonian

 New Zealand

A species of Symphysanodon.

Thalasseleotris whatua[36]

Sp. nov

Valid

Schwarzhans

Lillburnian

 New Zealand

A species of Thalasseleotris.

Tharsis elleri[80]

Sp. nov

Valid

Arratia, Schultze & Tischlinger

Late Jurassic (Tithonian)

Altmühltal Formation

 Germany

A teleost belonging to the family Ascalaboidae.

Thiollierepycnodus[81]

Gen. et comb. nov

Valid

Ebert

Late Jurassic

 France
 Germany

A member of the family Pycnodontidae; a new genus for "Pycnodus wagneri Thiollière (1852).

Thryptodus loomisi[82]

Sp. nov

Valid

Shimada

Late Cretaceous (CenomanianTuronian)

Britton Formation

 United States
( Texas)

A member of the family Plethodidae.

Tonganago wharenga[36]

Sp. nov

Valid

Schwarzhans

Waipawan to Bortonian

 New Zealand

A member of the family Congridae.

Tongarewa[36]

Gen. et 2 sp. nov

Valid

Schwarzhans

Duntroonian to Altonian

 New Zealand

A threefin blenny. The type species is T. waihaoensis; genus also includes T. clementsi.

Toroatherina[36]

Gen. et sp. et comb. nov

Valid

Schwarzhans

Eocene

 New Zealand
 United States

An Old World silverside. The type species is T. toroa; genus also includes "Otolithus (Mugilidarum)" debilis Koken (1891).

Toxopyge[60]

Gen. et sp. et comb. nov

Valid

Bradić-Milinović, Ahnelt & Schwarzhans in Bradić-Milinović et al.

Early Miocene

 Serbia
  Switzerland

A member of the family Gobiidae. The type species is T. campylus; genus also includes "Gobius" longus Salis (1967).

Trachyrincus aulax[36]

Sp. nov

Valid

Schwarzhans

Otaian to Tongaporutuan

 New Zealand

A species of Trachyrincus.

Tranawuen[58]

Gen. et comb. nov

Valid

Gouiric-Cavalli, Remírez & Kriwet

Early Cretaceous (Valanginian–early Hauterivian)

Agrio Formation

 Argentina

A member of Pycnodontiformes. Genus includes "Macromesodon" agrioensis.

Trawdenia[83]

Gen. et comb. nov

Valid

Coates & Tietjen

Carboniferous (Pennsylvanian)

 United Kingdom

An early ray-finned fish. The type species is "Rhadinichthys" planti Traquair (1888); genus also includes "Mesopoma" carricki Coates (1993) and "Mesopoma" pancheni Coates (1993).

Tunisiaclupea[84]

Gen. et sp. nov

Valid

Boukhalfa et al.

Early Cretaceous (Barremian)

Chotts Basin

 Tunisia

A member of Clupeomorpha belonging to the group Ellimmichthyiformes and to the family Paraclupeidae. Genus includes new species T. speratus.

Uranoscopus rudis[36]

Sp. nov

Valid

Schwarzhans

Duntroonian

Chatton Formation

 New Zealand

A species of Uranoscopus.

Uranoscopus tectiformis[36]

Sp. nov

Valid

Schwarzhans

Altonian

 New Zealand

A species of Uranoscopus.

Valenciennellus fastigatus[36]

Sp. nov

Valid

Schwarzhans

Otaian and Altonian

 New Zealand

A species of Valenciennellus.

Veridagon[85][86]

Gen. et sp. nov

Valid

Díaz-Cruz, Alvarado-Ortega & Carbot-Chanona

Late Cretaceous (Cenomanian)

Cintalapa Formation

 Mexico

A member of Aulopiformes belonging to the family Enchodontidae. Genus includes new species V. avendanoi.

Vinciguerria orientalis[87]

Sp. nov

Valid

Nam, Ko & Nazarkin

Middle Miocene

Duho Formation

 South Korea

A species of Vinciguerria.

Waihaoclupea[36]

Gen. et sp. nov

Valid

Schwarzhans

Mangaorapan to Bortonian

 New Zealand

A member of the family Clupeidae. The type species is W. pinguis.

Waitahana[36]

Gen. et comb. nov

Valid

Schwarzhans

Otaian to Waiauan

 New Zealand

A southern sandfish. The type species is "Trachinoideorum" sagittiformis Schwarzhans (1980); genus also includes "Citharus" latisulcatus Frost (1924) and "Trachinoideorum" ultimus Schwarzhans (1980).

Waitakia aho[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan to Porangan

 New Zealand

A member of the family Hemerocoetidae.

Waitakia proclinens[36]

Sp. nov

Valid

Schwarzhans

Mangaorapan to Bortonian

 New Zealand

A member of the family Hemerocoetidae.

Waitakia profunda[36]

Sp. nov

Valid

Schwarzhans

Kaiatan to Runangan

 New Zealand

A member of the family Hemerocoetidae.

Xenocephalus otaianus[36]

Sp. nov

Valid

Schwarzhans

Otaian

 New Zealand

A species of Xenocephalus.

Yudaiichthys[62]

Gen. et sp. nov

Valid

Xu et al.

Middle Triassic (Anisian)

Guanling Formation

 China

A member of Ginglymodi assigned to the group Kyphosichthyiformes and to the new family Lashanichthyidae. The type species is Y. eximius.

Zandtfuro[77]

Gen. et sp. nov

Valid

Ebert

Late Jurassic

 Germany

A member of Halecomorphi. Genus includes new species Z. tischlingeri.

Zonobythites cornifer[36]

Sp. nov

Valid

Schwarzhans

Porangan and Bortonian

 New Zealand

A member of the family Ophidiidae.

Lobe-finned fishes

Name Novelty Status Authors Age Type locality Country Notes Images

Anchidipterus[88]

Gen. et sp. nov

Valid

Krupina in Lebedev, Krupina & Linkevich

Devonian (Famennian)

 Russia
( Tver Oblast)

A lungfish. Genus includes new species A. dariae.

Clackodus[89]

Gen. et comb. nov

Valid

Smithson, Challands & Smithson

Carboniferous (Serpukhovian)

Clackmannan Group

 United Kingdom

A lungfish; a new genus for "Ctenodus" angustulus Traquair.

Ctenodus boudariensis[90]

Sp. nov

Valid

Kemp

Carboniferous (Viséan)

Ducabrook Formation

 Australia

A lungfish.

Isityumzi[91]

Gen. et sp. nov

Valid

Gess & Clement

Devonian (Famennian)

Witpoort Formation

 South Africa

A lungfish. The type species is I. mlomomde.

Limanichthys[92]

Gen. et sp. nov

Valid

Challands et al.

Carboniferous (Tournaisian)

Ballagan Formation

 United Kingdom

A lungfish. Genus includes new species L. fraseri.

Rubrognathus[93]

Gen. et sp. nov

Valid

Lebedev & Clément

Devonian (late Givetian–early Frasnian)

Yam-Tesovo Formation

 Russia
( Leningrad Oblast)

A member of Elpistostegalia. The type species is R. kuleshovi.

Selenodus[94]

Gen. et sp. nov

Valid

Mondéjar-Fernández

Devonian (Eifelian)

 Morocco

An onychodont sarcopterygian. Genus includes new species S. aquesbiae.

Whiteia uyenoteruyai[95]

Sp. nov

Valid

Yabumoto et al.

Early Triassic

Middle Sakamena Group

 Madagascar

Other fishes

Name Novelty Status Authors Age Type locality Country Notes Images

Aberrosquama[96]

Gen. et sp. nov

Valid

Burrow in Burrow et al.

Silurian (probably Gorstian)

Hamelin Formation

 Australia

A stem-gnathostome of uncertain phylogenetic placement. The type species is A. occidens.

General research

  • Revision of Early Devonian psammosteids described by Beverly Halstead from the so-called "Placoderm Sandstone" (Świętokrzyskie Mountains, Poland) is published by Dec (2019).[97]
  • A study on the locomotion of psammosteids, focusing on the influence of the shape of the psammosteid body form on hydrodynamic performance, is published by Dec (2019).[98]
  • A study on the anatomy of dermal plates of Astraspis is published by Lemierre & Germain (2019), who report possible evidence of presence of proliferative cartilage in this taxon.[99]
  • A study on the anatomy of the circulatory system of the head of Shuyu is published by Gai, Zhu & Donoghue (2019).[100]
  • Redescription of Sinogaleaspis shankouensis, based on data from 11 new specimens from the Silurian Xikeng Formation (Jiangxi, China), is published online by Gai et al. (2019).[101]
  • A study on the anatomy of the dermal skeleton of Tremataspis mammillata is published by O'Shea, Keating & Donoghue (2019).[102]
  • Redescription of Asterolepis orcadensis based on newly collected fossil material is published by Newman, den Blaauwen & Leather (2019).[103]
  • A study on the morphology of the jaw elements of a buchanosteoid placoderm specimen ANU V244 from the Early Devonian limestones (~400 Ma) at Burrinjuck, near Canberra (Australia) is published by Hu et al. (2019).[104]
  • A specimen of Dunkleosteus terrelli preserving vertebrae fused into a structure known as the synarcual is described from the Devonian Cleveland Shale Member of the Ohio Shale Formation (Ohio, United States) by Johanson et al. (2019).[105]
  • A redescription and a study on the phylogenetic relationships of a putative antiarch Silurolepis platydorsalis is published by Zhu, Lu & Zhu (2019), who reinterpret this species as a maxillate placoderm close to Qilinyu.[106]
  • New body fossils of Cheiracanthus intricatus, including the first known articulated specimen of this species, are described from the Givetian Tordalen Formation (Spitsbergen, Norway) by Newman, Burrow & den Blaauwen (2019).[107]
  • A study on the anatomy of the skeletal elements of the pharynx of Ptomacanthus anglicus, and on its implications for the knowledge of the evolution of the pharynx of jawed vertebrates, is published by Dearden, Stockey & Brazeau (2019).[108]
  • Description of fossils of Carboniferous (Mississippian) cartilaginous fishes from the area of Krzeszowice (Poland) is published by Ginter & Złotnik (2019).[109]
  • A study on microwear on teeth of Edestus minor, and on its implications for the knowledge of function of teeth of this fish, is published by Itano (2019).[110]
  • A revision of species belonging to the genus Edestus is published by Tapanila & Pruitt (2019).[111]
  • Discovery of a cast of the holotype of Petalodus ohioensis in the collections of the Yale Peabody Museum of Natural History is reported by Carpenter & Itano (2019), who consider the species Petalodus alleghaniensis to be a junior synonym of P. ohioensis.[112]
  • Restudy of a putative bill of an ibis-like bird from the Eocene La Meseta Formation (Antarctica) described by Jadwiszczak, Gaździcki & Tatur (2008)[113] is published by Agnolin, Bogan & Rozadilla (2019), who consider this specimen to be more likely to be a dorsal spine of a chimaeroid cartilaginous fish.[114]
  • A study on the anatomy of the skull of Tristychius arcuatus, providing evidence of adaptations for suction feeding, is published by Coates et al. (2019).[115]
  • Description of the first skeletal remains of Phoebodus from the Famennian of the Maïder region of Morocco, providing new information on the anatomy of this species, and a study on the phylogenetic affinities of Phoebodus is published by Frey et al. (2019).[116]
  • Teeth of a hybodont shark belonging to the genus Asteracanthus, with anatomy indicative of a crushing feeding behaviour, are described from the Upper Jurassic deposits of the Monte Nerone Pelagic Carbonate Platform, in the Umbria‐Marche‐Sabina Palaeogeographic Domain (Italy) by Citton et al. (2019).[117]
  • An association of 58 teeth of Ptychodus anonymus, representing the first occurrence of an associated dentition of this species, is described from the Cenomanian Jetmore Member of the Greenhorn Formation (Kansas, United States) by Hamm (2019).[118]
  • Description of new associated skeletal remains of Ischyrhiza mira from the Upper Cretaceous of Tennessee and Alabama and a study on the paleobiology of this species is published by Sternes & Shimada (2019).[119]
  • A study on the anatomy and phylogenetic relationships of Promyliobatis gazolai is published by Marramà et al. (2019).[120]
  • Fossils of members of the genus Aetomylaeus are described from localities in Peru and Chile by Villafaña et al. (2019), representing the first unambiguous fossil record of this genus from the Neogene of the southeastern Pacific.[121]
  • A specimen of the whiptail stingray species Tethytrygon muricatus preserving the uterus bearing four eggs is described from the Eocene of the Monte Bolca locality (Italy) by Fanti, Mazzuferi & Marramà (2019).[122]
  • Description of Langhian ray fossils from the Lower Tagus Basin (Portugal) is published by Fialho, Balbino & Antues (2019).[123]
  • A study on the taxonomic status and geological age of large shark remains from the Upper Cretaceous of the Castellavazzo locality (Italy) discovered in the 19th century is published by Conte et al. (2019).[124]
  • 15 partial skeletons of lamniform sharks, including the largest specimen of Cretoxyrhina mantelli known to date, are described from the Upper Cretaceous Scaglia Rossa Formation (Italy) by Amalfitano et al. (2019), who also review the taxonomic history of C. mantelli.[125]
  • A study on teeth histology and mineralization pattern in lamniform sharks, based on data from extant and fossil taxa (including enigmatic galeomorph shark Palaeocarcharias stromeri), is published by Jambura et al. (2019).[126]
  • A study on changes of diversity of lamniform sharks throughout their evolutionary history, aiming to determine the causes of their decline in the last 20 million years, is published by Condamine, Romieu & Guinot (2019).[127]
  • A study on changes in the presence or absence of lateral cusplets on teeth of members of the genus Carcharocles from the Calvert, Choptank, and St. Marys formations, and on their implications for the transition of shark populations from these formations from a Carcharocles chubutensis-dominated population to one dominated by C. megalodon, is published by Perez et al. (2019).[128]
  • A revision and a reevaluation of the reliability of all post-Messinian occurrences of Otodus megalodon in marine strata from western North America, and a study on the timing of extinction of this species, is published by Boessenecker et al. (2019).[129]
  • A study on the body size of Otodus megalodon, as inferred from the relationship between the ontogenetic development of teeth and total body length in the great white shark, will be published by Shimada (2019).[130]
  • Partial forelimb of a rorqual with several shark bite marks is described from the Pliocene Burica Formation (Panama) by Cortés et al. (2019).[131]
  • A study on the anatomy of the holotype specimen of a putative Paleocene shark Platyacrodus unicus is published by Bogan, Agnolin & Ezcurra (2019), who reinterpret this specimen as a carapace of a small retroplumid crab belonging to the genus Costacopluma.[132]
  • An assemblage of well-preserved isolated teeth of elasmobranchs is described from the late Oligocene of the North Alpine Foreland Basin (Austria) by Feichtinger et al. (2019).[133]
  • Description of the deep-sea elasmobranch fauna from the Miocene Yatsuo Group in central Japan, including the first fossil occurrences of the genera Arhynchobatis and Pseudoraja, will be published by Nishimatsu & Ujihara (2019).[134]
  • Description of the deep-sea elasmobranch fossils from the Miocene Makino Formation in southwest Japan, including the first fossil occurrences of the genera Springeria and Narke, is published by Nishimatsu (2019).[135]
  • A study on Paleocene cartilaginous fish fossils from the Lower Clayton Limestone Unit of the Midway Group near Malvern, Arkansas, evaluating the implications of these fossils for the knowledge of cartilaginous fish diversity across the Cretaceous-Paleogene boundary in the Malvern region and Gulf Coastal Plain of southwestern Arkansas, is published online by Maisch, Becker & Griffiths (2019).[136]
  • A study on the morphology of scales and squamation pattern in Guiyu oneiros is published by Cui, Qiao & Zhu (2019).[137]
  • A study on the anatomy and phylogenetic relationships of Brazilichthys macrognathus is published by Figueroa, Friedman & Gallo (2019).[138]
  • Permian species Palaeothrissum inaequilobum Blainville (1818) and P. parvum Blainville (1818) are found to be senior synonyms of the widely used species name Aeduella blainvillei (Agassiz, 1833) by Brignon (2019). Conditions exist for reversal of precedence and Aeduella blainvillei is declared nomen protectum.[139]
  • A study on the anatomy of the jaws, palate and teeth of Eurynotus crenatus is published by Friedman et al. (2019).[140]
  • New specimen of Birgeria liui, representing the most complete and articulated postcranial skeleton of an adult specimen of Birgeria reported so far, is described from the Ladinian of South China by Ni et al. (2019).[141]
  • Fossil remains of a spiral valve are reported in a specimen of Peipiaosteus pani from the Lower Cretaceous Yixian Formation (China) by Capasso (2019).[142]
  • Teeth of members of the genus Pycnodus are described from the Upper Cretaceous Cap de Naze marine formation (Senegal) by Capasso (2019).[143]
  • Description of a partial skeleton of Micropycnodon kansasensis from the Smoky Hill Chalk Member of the Niobrara Chalk (Kansas, United States), providing new information on the anatomy of this species, is published by Cronin & Shimada (2019).[144]
  • A study on the paleobiology of specimens of Nursallia gutturosum from the Cenomanian-Turonian platy limestone deposit of Vallecillo (north-eastern Mexico) is published by Stinnesbeck, Rust & Herder (2019).[145]
  • A study on the anatomy and phylogenetic relationships of Robustichthys luopingensis is published by Xu (2019).[146]
  • Fossil teeth representing the first evidence of Late Jurassic ginglymodians from Mt. Nerone in the Umbria-Marche-Sabina Domain (Italy) are described by Romano et al. (2019).[147]
  • A study on the anatomy and phylogenetic relationships of "Lepidotes" bernissartensis is published online by Cavin, Deesri & Olive (2019), who transfer this species to the genus Scheenstia.[148]
  • A study on the internal anatomy of a left jaw of a member of the genus Scheenstia from the Kimmeridgian Reuchenette Formation (Switzerland) is published online by Leuzinger et al. (2019), who describe a peculiar tooth replacement mode in this specimen.[149]
  • A study on the stomach contents of two specimens of Lepidotes from the Lower Jurassic of Germany will be published by Thies, Stevens & Stumpf (2019).[150]
  • Six new occurrences of Belonostomus, documenting the biogeographic and biostratigraphic range of this genus in North America, are reported from the Upper Cretaceous of Texas, Alabama and Mississippi by Van Vranken, Fielitz & Ebersole (2019).[151]
  • A study on the diversity of pectoral fin shape amongst members of Pachycormiformes is published by Liston et al. (2019).[152]
  • A study on the anatomy of the skull of Martillichthys renwickae will be published by Dobson et al. (2019).[153]
  • Fossil remains of a member or a relative of the genus Asthenocormus are described from the Upper Jurassic of the Ameghino (= Nordenskjöld) Formation of the Antarctic Peninsula by Gouiric-Cavalli et al. (2019), representing the first record of a suspension-feeding pachycormid from the Upper Jurassic of the Antarctic Peninsula and the oldest pachycormid yet recovered from Antarctica.[154]
  • New, three-dimensionally preserved specimens of Pachycormus are described from the Toarcian of Strawberry Bank at Ilminster (Somerset, United Kingdom) by Cawley et al. (2019).[155]
  • A study on the anatomy of Pleuropholis decastroi from the Lower Cretaceous (Albian) limestones of Pietraroja (Province of Benevento, Italy) and on the phylogenetic relationships of the family Pleuropholidae is published by Taverne & Capasso (2019).[156]
  • A study on the anatomy and phylogenetic relationships of the teleost species Majokia brasseuri from the Middle Jurassic Stanleyville Formation (Democratic Republic of the Congo) is published by Taverne (2019), who names a new order Majokiiformes.[157]
  • A study on the anatomy, jaw mechanics and phylogenetic relationships of Dugaldia emmilta is published by Cavin & Berrell (2019).[158]
  • A juvenile specimen of Xiphactinus audax, representing the smallest specimen of this species reported so far, will be described from the Cretaceous Niobrara Chalk (Kansas, United States) by King & Super (2019).[159]
  • A study on the internal structure of a fossil specimen of Notelops brama, investigated by neutron tomography, is published by Pugliesi et al. (2019).[160]
  • A study on the anatomy and phylogenetic relationships of Cavenderichthys talbragarensis and Waldmanichthys koonwarri is published online by Bean & Arratia (2019).[161]
  • A study on fossils of members of the genus Capoeta from the Pliocene locality Çevirme (Turkey) and on the evolutionary history of this genus is published by Ayvazyan, Vasilyan & Böhme (2019).[162]
  • Evidence of presence of managed aquaculture of the common carp by around 6000 BC is reported from the Early Neolithic Jiahu site (China) by Nakajima et al. (2019).[163]
  • A study on the morphology of fossil catfish spines from the Upper Cretaceous Adamantina and Marilia formations (Brazil) is published by Alves, Bergqvist & Brito (2019).[164]
  • A study on positions, heading directions and possible behavioural rules used in a group of fossil specimens of Erismatopterus levatus from the Eocene Green River Formation is published by Mizumoto, Miyata & Pratt (2019).[165]
  • Description of gadiform fossils from the Eocene sediments of the Sverdlovsk and Tyumen regions (Ural and Western Siberia, Russia), including fossils of members of the family Merlucciidae, is published by Marramà et al. (2019).[166]
  • Description of an incomplete percomorph specimen from the Miocene Ixtapa Formation (Mexico), representing the oldest primary freshwater percomorph fish from Mexico reported so far, is published by Cantalice & Alvarado-Ortega (2019).[167]
  • A study on the evolutionary history of the fish clade Pelagiaria is published by Friedman et al. (2019).[168]
  • A revision of the nomenclature of extant and fossil barracudas is published by Ballen (2019).[169]
  • A review of the published fossil record of the family Labridae is published by Bellwood et al. (2019).[170]
  • A study on the feeding habits of the percomorph fish Rhenanoperca minuta and other fishes from the Eocene Messel pit (Germany) is published by Micklich, Baranov & Wappler (2019).[171]
  • A study on the phylogenetic relationships of "psarolepid" bony fishes, evaluating which characters cause their different placements in analyses utilizing different methods for reconstructing the tree of life, is published by King (2019).[172]
  • A study on the ontogeny of the neurocranium and brain in the West Indian Ocean coelacanth, and on its implications for the knowledge of the evolution of the head of lobe-finned fishes, is published by Dutel et al. (2019).[173]
  • Redescription of Axelrodichthys araripensis and a comparative study of several other members of Mawsoniidae is published by Fragoso, Brito & Yabumoto (2019), who transfer the species Mawsonia lavocati to the genus Axelrodichthys.[174]
  • A study on the phylogenetic relationships and evolutionary history of mawsoniid coelacanths is published by Cavin et al. (2019).[175]
  • A study on the skull anatomy of Arquatichthys porosus, focusing on a newly-discovered postparietal shield, is published by Lu & Zhu (2019).[176]
  • Description of the posterior part of the skull of Tungsenia paradoxa is published by Lu et al. (2019).[177]
  • New fossil material of "Holoptychius" radiatus Newberry (1889) from the Devonian (Famennian) Catskill Formation (Pennsylvania, United States), providing new information on the anatomy of this species, is described by Daeschler, Downs & Matzko (2019), who transfer this species to the tristichopterid genus Langlieria.[178]
  • A study on the bone histology of the humerus of Hyneria lindae is published by Kamska et al. (2019).[179]
  • Description of new fossil material of Edenopteron from the Devonian (Famennian) Worange Point Formation (Australia) and a study on the phylogenetic relationships of this taxon is published online by Young et al. (2019).[180]
  • A study on the anatomy of the shoulder girdle and opercular series of Gogonasus andrewsae is published by Hu, Young & Lu (2019).[181]
  • A historical review of the fossil record of Devonian tetrapods and basal tetrapodomorphs from East Gondwana (Australasia, Antarctica) is published by Long, Clement & Choo (2019), who also present preliminary findings on the anatomy of the canowindrid Koharalepis jarviki based on synchrotron scan data.[182]
  • A study on the anatomy of fossil coelacanth lungs, on accessory air-breathing structures in fossil fishes and stem-tetrapods, and on the evolution of air breathing is published by Cupello, Clément & Brito (2019).[183]
  • A study on patterns of tooth replacement in Onychodus jandemarrai, Eusthenopteron foordi, Tiktaalik roseae and in extant West Indian Ocean coelacanth is published by Doeland et al. (2019).[184]
  • A study on changes of the skeletal anatomy of the pelvic and pectoral appendages during the transition from fins to limbs in vertebrate evolution, as indicated by data from fossil lobe-finned fishes and early tetrapods, is published by Esteve-Altava et al. (2019).[185]
  • A study on the anatomy of dermal rays in pectoral fins of Sauripterus taylori, Eusthenopteron foordi and Tiktaalik roseae, evaluating its implications for the knowledge of the evolution of dermal rays in early members of Tetrapodomorpha prior to the origin of digits, is published online by Stewart et al. (2019).[186]
  • A study on the evolution of the branchiostegal ray series in the skull of bony fishes, as indicated by data from extant and fossil taxa, is published by Ascarrunz et al. (2019).[187]
  • A study on the origin and evolution of acellular bone (bone without osteocytes) in fossil and extant actinopterygian fishes is published by Davesne et al. (2019).[188]
  • A diverse fish assemblage is reported from the Carboniferous (upper Pennsylvanian) Horquilla Formation (New Mexico, United States) by Ivanov & Lucas (2019).[189]
  • A study on the diversity and ecology of Triassic fish assemblages from the Villány Hills (Hungary) is published by Szabó, Botfalvai & Osi (2019).[190]
  • Cretaceous (probably BarremianAptian) fish fossils with strong affinities with Early Cretaceous faunas of Thailand are described from Pahang (Malaysia) by Teng et al. (2019).[191]
  • Description of a freshwater fish assemblage from the Cretaceous (Albian-Cenomanian) Açu Formation (Brazil) is published by Veiga, Bergqvist & Brito (2019).[192]
  • Description of Eocene (Bartonian) fish fauna from the Luna de Sus locality (Romania) is published by Trif, Codrea & Arghiuș (2019).[193]
  • A study on the composition of the otolith assemblage from the Santa Barbara Basin near the coast of California over the preceding two millennia is published by Jones & Checkley (2019).[194]
  • A study on the evolution of the herbivorous coral reef fishes, as indicated by data from extant and fossil species, is published by Siqueira, Bellwood & Cowman (2019).[195]

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