1994 in paleontology
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 1994.
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Wikimedia Commons has media related to 1994 in paleontology. |
Plants
Conifers
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen et sp nov |
valid |
Manchester |
Fossil Yew Genus, type species D. torreyoides |
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Sp nov |
valid |
Manchester |
Fossil Yew species |
|||||
Sp nov |
valid |
Manchester |
Fossil nutmeg yew species |
Angiosperms
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp nov |
valid |
Manchester |
A Kiwi species |
|||||
Alangium eydei[2] |
Sp nov |
valid |
Manchester |
A dogwood species |
||||
Alangium rotundicarpum[2] |
Sp nov |
valid |
Manchester |
A dogwood species |
||||
Anamirta leiocarpa[2] |
Sp nov |
valid |
Manchester |
A moonseed species |
||||
Anonaspermum bonesii[2] |
Sp nov |
valid |
Manchester |
A custard apple relative |
||||
Anonaspermum rotundum[2] |
Sp nov |
valid |
Manchester |
A custard apple relative |
||||
Atriaecarpum clarnense[2] |
Sp nov |
valid |
Manchester |
A moonseed species |
||||
Bursericarpum oregonense[2] |
Sp nov |
valid |
Manchester |
A torchwood family species |
||||
Calycocarpum crassicrustae[2] |
Sp nov |
valid |
Manchester |
A moonseed species |
||||
Castanopsis crepetii[2] |
Sp nov |
valid |
Manchester |
A chinkapin species |
||||
Comicilabium[2] |
Gen et sp nov |
valid |
Manchester |
An icacinaceous relative. |
||||
Sp nov |
valid |
Manchester |
A dogwood species |
|||||
Gen et sp nov |
valid |
Manchester |
A hazelnut relative, Type species C. hancockii |
|||||
Curvitinospora[2] |
Gen et sp nov |
valid |
Manchester |
A moonseed relative. |
||||
Davisicarpum limacioides[2] |
Sp nov |
valid |
Manchester |
A moonseed species. |
||||
Deviacer[2] |
Gen et sp nov |
valid |
Manchester |
|||||
Diploclisia auriformis[2] |
Comb nov |
jr synonym |
(Hollick) Manchester |
A moonseed species. |
||||
Emmenopterys dilcheri[2] |
Sp nov |
valid |
Manchester |
A rubiaceous species. |
||||
Eohypserpa scottii[2] |
Sp nov |
valid |
Manchester |
A moonseed species. |
||||
Sp nov |
valid |
Magallón-Puebl |
Pié de Vaca Formation |
Youngest Eucommia sp. in North America. |
||||
Fortunearites[2] |
Gen et sp nov |
valid |
Manchester |
A witch-hazel relative, type species F. endressii |
||||
Hydrangea knowltonii[2] |
Sp nov |
valid |
Manchester |
A Hydrangea species. |
||||
lodes chandlerae[2] |
Sp nov |
valid |
Manchester |
An icacinaceous species. |
||||
Iodicarpa[2] |
Gen et sp nov |
valid |
Manchester |
An icacinaceous relative. |
||||
Gen et sp nov |
valid |
Manchester |
A hazelnut relative, type species K. parvum |
|||||
Lindera clarnensis[2] |
Sp nov |
valid |
Manchester |
|||||
Laurocalyx wheelerae[2] |
Sp nov |
valid |
Manchester |
|||||
Laurocarpum hancockii[2] |
Sp nov |
valid |
Manchester |
|||||
Laurocarpum nutbedensis[2] |
Sp nov |
valid |
Manchester |
|||||
Laurocarpum raisinoides[2] |
Sp nov |
valid |
Manchester |
|||||
Magnolia muldoonae[2] |
Sp nov |
valid |
Manchester |
A Magnolia species |
||||
Magnolia paroblonga[2] |
Sp nov |
valid |
Manchester |
A Magnolia species |
||||
Magnolia tiffneyi[2] |
Sp nov |
valid |
Manchester |
A Magnolia species |
||||
Mastixicarpum occidentale[2] |
Sp nov |
valid |
Manchester |
A Tupelo relative |
||||
sp nov |
valid |
Manchester |
Middle Eocene |
Clarno Formation |
A sabiaceous species. |
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Meliosma bonesii[2] |
sp nov |
valid |
Manchester |
Middle Eocene |
Clarno Formation |
A sabiaceous species. |
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Meliosma elongicarpa[2] |
sp nov |
valid |
Manchester |
Middle Eocene |
Clarno Formation |
A sabiaceous species. |
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Meliosma leptocarpa[2] |
sp nov |
valid |
Manchester |
Middle Eocene |
Clarno Formation |
A sabiaceous species. |
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Nyssa scottii[2] |
sp nov |
valid |
Manchester |
Middle Eocene |
Clarno Formation |
A Tupelo relative |
||
Comb nov |
valid |
(Scott) Manchester |
Middle Eocene |
Clarno Formation |
||||
Palaeoallophylus gordonii[2] |
Sp nov |
valid |
Manchester |
A soapberry species. |
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Palaeoallophylus gordonii[2] |
Sp nov |
valid |
Manchester |
A soapberry species. |
||||
Gen et sp nov |
valid |
Manchester |
A ginseng genus, type species P. oregonensis |
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Paleoplatycarya? hickeyi[2] |
Sp nov |
valid |
Manchester |
A Platycarya relative species. |
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Pentoperculum[2] |
Gen et comb nov |
valid |
Manchester |
A Sumac relative. |
||||
Platanus hirticarpa[2] |
Sp nov |
valid |
Manchester |
A platanaceous Infructescence species. |
||||
Prunus olsonii[2] |
Sp nov |
valid |
Manchester |
An cherry/plum species. |
||||
Prunus weinsteinii[2] |
Sp nov |
valid |
Manchester |
An cherry/plum species. |
||||
Pyrenacantha occidentalis[2] |
Sp nov |
valid |
Manchester |
An icacinaceous species. |
||||
Sp nov |
valid |
Manchester |
A Sumac species |
|||||
Sabia prefoetida[2] |
Comb nov |
valid |
(Becker) Manchester |
Late Eocene |
Ruby flora |
A sabiaceous species. |
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Bumelia? globosa[2] |
Sp nov |
jr synonym |
(Manchester) Manchester |
A lardizabalaceous species; |
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Saxifragispermum tetragonalis[2] |
Sp nov |
valid |
Manchester |
A willow relative |
||||
Tanyoplatanus[2] |
Gen et sp nov |
valid |
Manchester |
A platanaceous Infructescence genus. |
||||
'Tinomiscoidea occidentalis[2] |
Sp nov |
valid |
Manchester |
A moonseed species |
||||
'Tinospora elongata[2] |
Sp nov |
valid |
Manchester |
A moonseed species |
||||
Thanikaimonia[2] |
Gen et sp nov |
valid |
Manchester |
A moonseed genus. |
||||
'Tinospora hardmanae[2] |
Sp nov |
valid |
Manchester |
A moonseed species |
||||
Arthropods
Newly named insects
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp nov |
Valid |
Baroni Urbani & De Andrade |
first fossil Acanthognathus species |
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Aeshna andancensis[8] |
Sp nov |
Valid |
Nel & Brisac |
Montagne d'Andance |
An Aeshna dragonfly |
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Aeshna cerdanica[8] |
Sp nov |
Valid |
Nel & Martínez-Delclòs |
An Aeshna dragonfly |
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Aeschnophlebia miocenica[8] |
Sp nov |
Valid |
Nel & Escuillé |
Montagne d'Andance |
An Aeschnophlebia dragonfly |
| ||
Sp nov |
Valid |
De Andrade |
A ponerin ant |
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Sp nov |
Valid |
De Andrade |
A ponerin ant |
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Sp nov |
Valid |
De Andrade |
A ponerin ant |
| ||||
Sp nov |
Valid |
De Andrade |
A ponerin ant |
| ||||
Sp nov |
Valid |
De Andrade |
A ponerin ant |
| ||||
Sp nov |
Valid |
De Andrade |
A ponerin ant |
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Calopteryx andancensis |
Sp nov |
Valid |
Nel & Brisac |
Montagne d'Andance |
A Calopteryx damselfly |
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Gen et comb nov. |
Valid |
Bolton |
Taimyr amber |
A Sphecomyrminae ant. |
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Forficula paleoligura[11] |
Sp nov |
Valid |
Nel, Albouy, Caussanel, & Jamet |
An earwig |
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Sp nov. |
Valid |
De Andrade |
Burdigalian |
Dominican Amber |
A myrmicin ant |
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Sp nov. |
Valid |
De Andrade |
Burdigalian |
Dominican Amber |
A myrmicin ant |
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Protoaristenymphes[12] |
Gen et sp nov |
Valid |
Nel & Henrotay |
A Mesochrysopid lacewing, type species P. bascharagensis |
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Sympecma incerta[13] |
Comb. nov |
Valid |
Piton |
Lake Chambon |
A Lestid damselfly, |
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Molluscs
Bivalves
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
sp nov |
Valid |
Elicki |
Zwetau Formation |
early Cambrian bivalve species |
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Fishes
Newly named bony fishes
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
valid |
Schwimmer, Stewart & Williams |
late Santonian to mid-Campanian |
A latimeriid coelacanth.[16] |
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Archosauromorphs
- Aff. Rebbachisaurus gastroliths documented.[17]
- The "Talkeetna Mountains Hadrosaur" specimen was discovered in a quarry being excavated for road material.[18] The quarry is near the Glenn Highway, approximately 150 miles northeast of Anchorage.[19] That fall, excavation began, and was resumed in the summer of 1996.[18]
Newly named pseudosuchians
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen et sp nov |
Valid |
Hunt & Lucas |
Late Triassic (late Carnian) |
A relative of Revueltosaurus. |
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Gen et sp nov |
Hunt & Lucas |
Late Triassic (late Carnian) |
A junior synonym of Revueltosaurus. |
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Newly named basal dinosauromorphs
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen et comb nov |
Valid |
Sereno & Arcucci |
Middle Triassic (Ladinian) |
A basal dinosauriform; |
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Newly named dinosaurs
Data courtesy of George Olshevsky's dinosaur genera list.[22]
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen et sp nov |
Valid |
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Gen et comb nov |
Valid |
Chatterjee & Creisler |
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Gen et sp nov |
Valid |
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Cheng |
described as Huabeisaurus allocotus in 2000 |
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Gen et sp nov |
Valid |
Currie, Godfrey, & Nesov |
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Gen et sp nov |
Valid |
Hammer & Hickerson |
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Gen et sp nov |
Valid |
Chure |
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Gen et sp nov |
Valid |
Zhao & Currie |
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Gen et sp nov |
Valid |
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Gen et sp nov |
Valid |
Perez-Moreno, Sanz, Buscalioni, |
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Gen et sp nov |
Valid |
Martin, Buffetaut, & Suteethorn |
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Gen et sp nov |
Valid |
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Gen et sp nov |
Valid |
Currie & Zhao |
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Gen et sp nov |
Valid |
Rich & Vickers-Rich |
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“Tonouchisaurus” |
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Gen et sp nov |
Valid |
Zhou |
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Newly named birds
Name | Status | Novelty | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Valid |
Gen. nov. et Sp. nov. |
Late Paleocene |
Type species B. orbisantiqui |
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Valid |
Sp. nov. |
probably Pleistocene |
Cave deposits |
A Strigidae. |
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Falco duboisi [39] |
Valid |
Sp. nov. |
Graham S. Cowles |
Cave deposits |
A Falconidae. |
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Mascarenachen kervazoi [39] |
Valid |
Gen. nov. et Sp. nov. |
Graham S. Cowles |
An Anatidae, type species M. kervazoi; |
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Mascarenotus grucheti [40] |
Valid |
Sp. nov. |
Roger Bour François Moutou Sonia Ribes |
A Strigidae. |
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Messelastur gratulator [41] |
Valid |
Gen. nov. et Sp. nov. |
Dieter S. Peters |
Middle Eocene |
MN 11 |
A Psittaciformes, Messelasturidae G. Mayr, 2005, this is the type species of the new genus. |
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Valid |
Sp. nov. |
Gerard F. van Tets |
Peat Swamp near Mining Lease |
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Valid |
Gen. nov. et Sp. nov. |
Santonian-Coniacian, Rio Colorado Formation |
An Enantiornithes Walker, 1981, Avisauridae Brett-Surman et Paul, 1985, this is the type species of the new genus. |
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Nycticorax borbonensis [39] |
Not valid |
Sp. nov. |
Graham S. Cowles |
An Ardeidae, this is a junior synonym of Nycticorax duboisi (Rothschild, 1907). |
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Otogornis genghisi [44] |
Valid |
Gen. nov. et Sp. nov. |
Hou Lianhai |
An Enantiornithes Walker, 1981, Longirostravidae Zhou, 2006, this is the type species of the new genus. |
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Presbyornis isoni [45] |
Valid |
Sp. nov. |
Late Paleocene |
An Anseriformes, Presbyornithidae Wetmore, 1926. |
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Valid |
Sp. nov. |
Late Quaternary |
Cave Deposits, South Island |
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Valid |
Sp. nov. |
Holocene. (Subrecent) |
An Accipitridae. |
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Wanshuina lii [48] |
Valid |
Gen. nov. et Sp. nov. |
Hou Lianhai |
Doumu Formation |
A Rallidae, this is the type species of the new genus. |
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Plesiosaurs
- Plesiosaur gastroliths documented.[49]
New taxa
Name | Status | Authors | Notes | |
---|---|---|---|---|
Valid |
Chatterjee Creisler |
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Pterosaurs
New taxa
Name | Status | Authors | Age | Unit | Location | Notes | Images | |
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Valid |
Frey Martill |
Early Cretaceous (late Aptian/early Albian) |
A pteranodontoid pterosaur. |
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Valid |
Padian, K., Horner de Ricqlès, A.J. |
Late Cretaceous (Judithian) |
An azhdarchid from Montana. |
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nomen nudum |
Bakker |
Late Jurassic (Kimmeridgian-Tithonian) |
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Valid |
Cai, Z. Wei, F. |
Late Cretaceous (early Campanian) |
A Chinese azhdarchid. |
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Other diapsids
New taxa
Name | Status | Authors | Age | Unit | Location | Notes | |
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Lucianosaurus[20] | Valid Non-Dinosaurian Taxon |
|
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Late Triassic (Norian) |
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Tecovasaurus[20] | Valid Non-Dinosaurian Taxon |
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Late Triassic (late Carnian) |
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Footnotes
- Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
- Manchester, S.R. (1994). "Fruits and Seeds of the Middle Eocene Nut Beds Flora, Clarno Formation, Oregon". Palaeontographica Americana. 58: 30–31.
- Myers, J.; Erwin, D. (2015). "Deviacer pidemarmanii sp. nov. (Polygalaceae) from the Late Eocene–Early Oligocene Badger's Nose Paleoflora, Modoc County, California". International Journal of Plant Sciences. 176 (3): 259–268. doi:10.1086/679759.
- Han, M.; Manchester, S. R.; Fu, Q.-Y.; Jin, J.-H.; Quan (2018). "Paleogene fossil fruits of Stephania (Menispermaceae) from North America and East Asia". Journal of Systematics and Evolution. 56 (2): 81–91. doi:10.1111/jse.12288. Unknown parameter
|First5=
ignored (|first5=
suggested) (help) - Call, V.B.; Dilcher, D.L. (1997). "The fossil record of Eucommia (Eucommiaceae) in North America" (PDF). American Journal of Botany. 84 (6): 798–814. doi:10.2307/2445816. JSTOR 2445816. PMID 21708632.
- Manchester, S. R. (1999). "Biogeographical relationships of North American tertiary floras". Annals of the Missouri Botanical Garden. 86: 472–522.
- Baroni Urbani, C.; De Andrade, M. L. (1994). "First description of fossil Dacetini ants with a critical analysis of the current classification of the tribe (Amber Collection Stuttgart: Hymenoptera, Formicidae. VI: Dacetini)". Stuttgarter Beiträge zur Naturkunde. Serie B (Geologie und Paläontologie). 198: 1–65.
- Nel, A.; Martinez-Delclòs, X.; Escuillé, F; Brisac, P. (1994). "Names; Les Aeshnidae fossiles: Etat actuel des connaissances (Odonata, Anisoptera)". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 194 (2/3): 143–186.
- De Andrade, M. L. (1994). "Fossil Odontomachiti Ants from the Dominican Republic (Amber Collection Stuttgart: Hymenoptera, Formicidae. VII: Odontomachiti)". Stuttgarter Beiträge zur Naturkunde. Serie B (Geologie und Paläontologie). 199: 1–28.
- Grimaldi, D.; Agosti, D.; Carpenter, J. M. (1997). "New and rediscovered primitive ants (Hymenoptera, Formicidae) in Cretaceous amber from New Jersey, and their phylogenetic relationships". American Museum Novitates. 3208: 1–43.
- Nel, A.; Albouy, V.; Caussanel, C; Jamet, C. (1994). "Réflexion paléo-entomologique sur la systématique des Dermaptères. Quatre nouveaux forficules fossiles de l'Oligocène de Provence (France)". Bulletin de la Société Entomologique de France. 99: 253–266.
- Nel, A; Henrotay, M. (1994). "Les Chrysopidae Mésozoïques. État actuel des connaissances. Description d'un nouveau genre et nouvelle espèce dans le Jurassique inférieur (Lias) (Insecta: Neuroptera)". Annales de la Société Entomologique de France. 30: 295–318.
- Nel, A.; Paicheler, J.-C. (1994). "Les Lestoidea (Odonata, Zygoptera) fossiles : un inventaire critique". Annales de Paléontologie. 80 (1): 1–59.
- Elicki, O.; Gürsu, S. (2009). "First record of Pojetaia runnegari Jell, 1980 and Fordilla Barrande, 1881 from the Middle East (Taurus Mountains, Turkey) and critical review of Cambrian bivalves" (PDF). Paläontologische Zeitschrift. 83 (2): 267–291. doi:10.1007/s12542-009-0021-9.
- David R. Schwimmer; J. D. Stewart; G. Dent Williams (1994). "Giant fossil coelacanths of the Late Cretaceous in the eastern United States". Geology. 22 (6): 503–506. doi:10.1130/0091-7613(1994)022<0503:gfcotl>2.3.co;2.
- Hugo Dutel; John G. Maisey; David R. Schwimmer; Philippe Janvier; Marc Herbin; Gaël Clément (2012). "The Giant Cretaceous Coelacanth (Actinistia, Sarcopterygii) Megalocoelacanthus dobiei Schwimmer, Stewart & Williams, 1994, and Its Bearing on Latimerioidei Interrelationships". PLOS ONE. 7 (11): e49911. doi:10.1371/journal.pone.0049911. PMC 3507921. PMID 23209614.
- Calvo (1994). Sanders, Manley, and Carpenter (2001), "Table 12.1" page 167.
- "Introduction," in Pasche and May (2001); page 220.
- "Location and Geologic Setting," in Pasche and May (2001); page 220.
- Hunt, A.P. and S.G. Lucas. 1994. Ornithischian dinosaurs from the Upper Triassic of the United States. In: the Shadow of the Dinosaurs, Early Mesozoic Tetrapods (N.C. Fraser and H.-D. Sues, Eds.). Cambridge Univ. Press, Cambridge, UK: pp. 227-241.
- Sereno, P.C.; Arcucci, A.B. (1994). "Dinosaur precursors from the Middle Triassic of Argentina: Marasuchus lilloensis gen. nov". Journal of Vertebrate Paleontology. 14: 53–73. doi:10.1080/02724634.1994.10011538.
- Olshevsky, George. "Dinogeorge's Dinosaur Genera List". Retrieved 2008-08-07.
- Sereno, P.C., J.A. Wilson, H.C.E. Larsson, D.B. Dutheil, H.-D. Sues. 1994. Early Cretaceous dinosaurs from the Sahara. Science 266 (14 October [5183]): pp. 267-271.
- Chatterjee, S.; Creisler, B.S. (1994). "Alwalkeria (Theropoda) and Morturneria (Plesiosauria), new names for preoccupied Walkeria Chatterjee, 1987 and Turneria". Journal of Vertebrate Paleontology. 14: 142. doi:10.1080/02724634.1994.10011546.
- Russell, D.A. and Z.-M. Dong. 1994. The affinities of a new theropod from the Alxa Desert, Inner Mongolia, People's Republic of China. Can. J. Earth Sci. 30:pp. 2107-2127.
- Currie, P.J., S.J. Godfrey, and L. Nesov. 1993 [Olshevsky cites 1994]. New caenagnathid (Dinosauria: Theropoda) specimens from the Upper Cretaceous of North America and Asia. Can. J. Earth Sci. 30: pp. 2255-2272.
- Hammer, W.R.; Hickerson, W.J. (1994). "A Crested Theropod Dinosaur from Antarctica". Science. 264: 828–830. Bibcode:1994Sci...264..828H. doi:10.1126/science.264.5160.828. PMID 17794724.
- Chure, D.J. 1994. Koparion douglassi, a new dinosaur from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument; The oldest troodontid (Theropoda: Maniraptora). Brigham Young University Geology Studies 40:pp. 11-15.
- Zhao, X.J. and P.J. Currie, P.J. 1994 [not 1993]. A large crested theropod from the Jurassic of Xinjiang, China. Canadian. J. Earth Sci. 30: pp. 2027-2036.
- Kirkland, J.I. and K. Carpenter. 1994. North America's First Pre-Cretaceous Ankylosaur (Dinosauria) from the Upper Jurassic Morrison Formation of Western Colorado. Brigham Young University Geology Studies 40: pp. 25-42.
- Perez-Moreno, B.; Sanz, J.L.; Buscalloni, A.D.; Moratalla, J.J.; Ortega, F.; Rasskin-Gutman, D. (1994). "A unique multitoothed ornithomimosaur dinosaur from the Lower Cretaceous of Spain". Nature. 370: 363–367. doi:10.1038/370363a0.
- Martin, V. E. Buffetaut, and V. Suteethorn. 1994. A new genus of sauropod dinosaur from the Sao Khua Formation (Late Jurassic or Early Cretaceous) of north-eastern Thailand. Comptes Rendus l'Acad. Sci. Paris (II) 319 : pp. 1085-1092.
- Russell, D.A. and Z.-M Dong. 1993 [George Olshevsky cites 1994, not 1993]. A nearly complete skeleton of a new troodontid dinosaur from the Early Cretaceous of the Ordos Basin, Inner Mongolia, People's Republic of China. Can. J. Earth Sci. 30: pp. 2163-2173.
- Currie, P.J. and X.J. Zhao. 1994. A new carnosaur (Dinosauria, Theropoda) from the Jurassic of Xinjiang, People's Republic of China. Can. J. Earth Sci., 30: pp. 2037-2081.
- Rich, T.H., and P. Vickers-Rich. 1994. Neoceratopsians and ornithomimosaurs: Dinosaurs of Gondwana origin. Res. Exploration 10: pp. 129-131.
- Zhou Songlin, 1994. Record of a fossil stegosaur from Yingshan in the Sichuan Basin. Sichuan Cultural Relics. 1994-S1: 8-14.
- Cécile Mourer-Chauviré (1994). "A Large Owl from the Paleocene of France" (PDF). Palaeontology. 37 (2): 339–348.
- Oscar Arredondo; Storrs L. Olson (1994). "A New Species of Owl of the Genus Bubo from the Pleistocene of Cuba (Aves: Strigiformes)" (PDF). Proceedings of the Biological Society of Washington. 107 (3): 436–444. Archived from the original (PDF) on 2014-08-09. Retrieved 2014-08-09.
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