2016 in arthropod paleontology

This list of fossil arthropods described in 2016 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2016, as well as other significant discoveries and events related to arthropod paleontology that occurred in the year 2016.

List of years in arthropod paleontology
In paleontology
2013
2014
2015
2016
2017
2018
2019

Arachnids

Research

  • A study on the phylogenetic relationships of the Eocene tick Ixodes succineus is published by Dunlop et al. (2016).[1]
  • A study on the phylogenetic relationships of the Carboniferous species Archaeometa nephilina and Arachnometa tuberculata is published by Selden, Dunlop & Garwood (2016), who interpret these taxa as harvestmen rather than spiders.[2]

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Aenigmatarbus[3]

Gen. et sp. nov

Valid

Poschmann et al.

Carboniferous (late Gzhelian)

 France

A member of Trigonotarbida of uncertain phylogenetic placement. The type species is Aenigmatarbus rasteli.

Afronothrus ornosae[4]

Sp. nov

Valid

Arillo, Subías & Sánchez-García

Early Cretaceous

 Spain

An oribatid mite, a species of Afronothrus.

Archaeoananteroides[5]

Gen. et sp. nov

Valid

Lourenço & Velten

Cretaceous

Burmese amber

 Myanmar

A scorpion belonging to the family Buthidae. The type species is A. maderai.

Betaburmesebuthus bellus[6]

Sp. nov

Valid

Lourenço

Early Cretaceous

Burmese amber

 Myanmar

A scorpion belonging to the superfamily Buthoidea and the family Palaeoburmesebuthidae.

Betaburmesebuthus bellus

Betaburmesebuthus fleissneri[7]

Sp. nov

Valid

Lourenço & Velten

Cretaceous

Burmese amber

 Myanmar

A scorpion belonging to the superfamily Buthoidea and the family Palaeoburmesebuthidae.[6]

Betaburmesebuthus larafleissnerae[8]

Sp. nov

Valid

Lourenço & Velten

Cretaceous

Burmese amber

 Myanmar

A scorpion belonging to the superfamily Buthoidea and the family Palaeoburmesebuthidae.

Burmesescorpiops[9]

Gen. et sp. nov

Valid

Lourenço

Early Cretaceous

Burmese amber

 Myanmar

A scorpion belonging to the family Palaeoeuscorpiidae and the subfamily Archaeoscorpiopinae. The type species is Burmesescorpiops groehni.

Calyptostoma katyae[10]

Sp. nov

Valid

Konikiewicz, Wohltmann & Mąkol

Eocene

Baltic amber

Europe (Baltic Sea coast)

A mite belonging to the group Prostigmata, a species of Calyptostoma.

Carbolohmannia[11]

Gen. et sp. nov

Valid

Robin et al.

Carboniferous (ca. 320 mya)

 China

An oribatid mite belonging to the group Mixonomata. The type species is Carbolohmannia maimaiphilus.

Chaerilobuthus gigantosternum[9]

Sp. nov

Valid

Lourenço

Early Cretaceous

Burmese amber

 Myanmar

A scorpion belonging to the family Chaerilobuthidae; a species of Chaerilobuthus.

Chaerilobuthus serratus[9]

Sp. nov

Valid

Lourenço

Early Cretaceous

Burmese amber

 Myanmar

A scorpion belonging to the family Chaerilobuthidae; a species of Chaerilobuthus.

Cheletomimus (Hemicheyletia) crinitus[12]

Sp. nov

Valid

Bochkov & Sidorchuk

Eocene

Baltic amber

Baltic Sea coast, most likely  Russia (Kaliningrad Oblast)

A mite belonging to the family Cheyletidae, a species of Cheletomimus.

Electroblemma[13]

Gen. et sp. nov

Valid

Selden, Zhang & Ren

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A member of Tetrablemmidae. The type species is E. bifida.

Electrokoenenia[14]

Gen. et sp. nov

Valid

Engel & Huang in Engel et al.

Late Cretaceous (Cenomanian)

 Myanmar

A microwhip scorpion. The type species is Electrokoenenia yaksha.

Hypovertex hispanicus[4]

Sp. nov

Valid

Arillo, Subías & Sánchez-García

Early Cretaceous

 Spain

An oribatid mite, a species of Hypovertex.

Idmonarachne[15]

Gen. et sp. nov

Valid

Garwood et al.

Carboniferous (Gzhelian)

 France

A member of Arachnida of uncertain phylogenetic placement, probably closely related to spiders. The type species is Idmonarachne brasieri.

Kronocharon prendinii adornellae[16]

Subsp. nov

Valid

Rossi & Merendino

Cretaceous

Burmese amber

 Myanmar

A whip spider.

Nothrus vazquezae[4]

Sp. nov

Valid

Arillo, Subías & Sánchez-García

Early Cretaceous

 Spain

An oribatid mite, a species of Nothrus.

?Opsieobuthus tungeri[17]

Sp. nov

Valid

Dunlop et al.

Early Permian (Sakmarian)

Leukersdorf Formation

 Germany

A scorpion belonging to the family Centromachidae; possibly a species of Opsieobuthus.

Parageralinura marsiglioi[18]

Sp. nov

Valid

Selden, Dunlop & Simonetto

Carboniferous (KasimovianGzhelian)

Probably Pizzul Formation

 Italy

A member of Thelyphonida, a species of Parageralinura.

Paraliochthonius miomaya[19]

Sp. nov

Valid

Judson

Miocene

 Mexico

A pseudoscorpion belonging to the family Chthoniidae; a species of Paraliochthonius.

Petrobunoides[20]

Gen. et sp. nov

Valid

Selden et al.

Late Cretaceous (early Cenomanian)

Burmese amber

 Myanmar

A member of Opiliones belonging to the family Epedanidae. The type species is Petrobunoides sharmai.

Platyliodes sellnicki[4]

Sp. nov

Valid

Arillo, Subías & Sánchez-García

Early Cretaceous

 Spain

An oribatid mite, a species of Platyliodes.

Porttrombidium gedanense[21]

Sp. nov

Valid

Konikiewicz, Sontag & Mąkol

Eocene (Lutetian)

Baltic amber

 Russia (Kaliningrad Oblast)

A mite belonging to the group Parasitengona and the family Microtrombidiidae.

Protofeaella[22]

Gen. et sp. nov

Valid

Henderickx in Henderickx & Boone

Late Cretaceous (Cenomanian)

Burmese amber

 Myanmar

A pseudoscorpion. Originally classified as a member of the family Feaellidae;[22] Wolfe et al. (2016) only considered it to be a likely member of the pseudoscorpion crown group, probably a feaelloid.[23] The type species is P. peetersae.

Rhopalurus renelauerae[24]

Sp. nov

Valid

Lourenço in Lourenço & Velten

Miocene

Dominican amber

 Dominican Republic

A scorpion belonging to the family Buthidae, a species of Rhopalurus.

Tenuelamellarea estefaniae[4]

Sp. nov

Valid

Arillo, Subías & Sánchez-García

Early Cretaceous

 Spain

An oribatid mite, a species of Tenuelamellarea.

Crustaceans

Research

Malacostracans

Name Novelty Status Authors Age Unit Location Notes Images

Acanthodromia zannatoi[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Dynomenidae.

Acanthogalathea broglioi[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A member of Galatheidae.

Acanthogalathea devecchii[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A member of Galatheidae.

Acanthogalathea paucispinosa[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A member of Galatheidae.

Aetocarcinus[33]

Gen. et comb. et sp. nov

Valid

Schweitzer et al.

Early Cretaceous (Albian)

Glen Rose Formation

 United States

A member of Raninoida belonging to the family Orithopsidae. The type species is "Diaulax" roddai Bishop (1983); genus also includes new species Aetocarcinus muricatus.

Alponella[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Goneplacoidea and the family Euryplacidae. The type species is A. paleogenica.

Amekicarcinus[34]

Gen. et sp. nov

Valid

Schweitzer, Odumodu & Feldmann

Eocene

 Nigeria

A crab. Genus includes new species A. enigmaticus.

Arcantitanais[35]

Gen. et sp. nov

Valid

Sánchez-García et al.

Cretaceous (AlbianCenomanian)

Charentese amber

 France

A member of Tanaidacea of uncertain phylogenetic placement. The type species is A. turpis.

Arcticocarcinus[36]

Gen. et comb. nov

Valid

Schweitzer et al.

Paleocene (Danian)

 Denmark

A crab belonging to the group Raninoida and the family Necrocarcinidae. A new genus for "Necrocarcinus" insignis Segerberg (1900).

Armadillopsis[35]

Gen. et sp. nov

Valid

Sánchez-García et al.

Late Cretaceous (Cenomanian)

Pyrenean amber

 France

A member of Tanaidacea of uncertain phylogenetic placement. The type species is A. rara.

Atherfieldastacus[37]

Gen. et comb. nov

Valid

Simpson in Robin et al.

Early Cretaceous

Atherfield Clay Formation

 Colombia[38]
 Germany
 Mexico
 South Africa
 Spain[39]
 United Kingdom

A member of Glypheoidea belonging to the family Mecochiridae. The type species is "Meyeria" magna M'Coy (1849); genus also includes Atherfieldastacus mexicanus (Rathbun, 1935), Atherfieldastacus rapax (Harbort, 1905) and Atherfieldastacus schwartzi (Kitchin,1908) .

Aulavescus paintenensis[40]

Sp. nov

Valid

Feldmann et al.

Late Jurassic

 Germany

A munidopsid galatheoid, a species of Aulavescus.

Barapseudia[41]

Gen. et sp. nov

Valid

Quayle

Eocene

Barton Clay Formation

 United Kingdom

A member of Tanaidacea belonging to the family Apseudidae. The type species is Barapseudia prima.

Basadromia[42]

Gen. et sp. nov

Valid

Artal et al.

Eocene (Priabonian)

 Spain

A member of Dromioidea, possibly a member of the family Dromiidae. The type species is Basadromia longifrons.

Bathynomus kominatoensis[43]

Sp. nov

Valid

Kato, Kurihara & Tokita

Late Miocene

Amatsu Formation

 Japan

A giant isopod.

Biohermia[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Homolodromioidea and the family Goniodromitidae. The type species is B. chalmasi.

Bittnereus depressus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Panopeidae.

Bittnereus tumidus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Panopeidae.

Bittnerilia granosa[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Parthenopidae.

Bolcagalathea[32]

Gen. et 3 sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A member of Galatheidae. The type species is B. corallina; genus also includes B. venetica and B. multispinosa.

Bolcapisa[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Epialtidae. The type species is B. giulianae.

Braggilambrus[45]

Gen. et sp. nov

Valid

De Angeli & Caporiondo

Eocene (Ypresian)

 Italy

A crab belonging to the family Parthenopidae. The type species is B. tani.

Branchioplax cordata[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Mathildellidae.

Bylgia anjobea[46]

Sp. nov

Valid

Winkler

Late Jurassic

Painten Formation

 Germany

A member of Penaeidae, a species of Bylgia.

Calappilia calculosa[47]

Sp. nov

Valid

Rumsey, Klompmaker & Portell

Late Eocene to early Oligocene

 United States

A member of Calappidae, a species of Calappilia.

Caporiondolus[48]

Gen. et sp. nov

Valid

De Angeli

Eocene (Priabonian)

 Italy

A crab belonging to the group Cyclodorippoida and the family Cymonomidae. The type species is Caporiondolus bericus.

Catillogalathea[49]

Gen. et 3 sp. nov

Valid

Robins et al.

Late Jurassic

 Austria
 Romania

A member of Galatheoidea. The type species is Catillogalathea falcula; genus also includes Catillogalathea patruliusi and Catillogalathea purcarenensis

Ceratiocaris winneshiekensis[50]

Sp. nov

Valid

Briggs et al.

Ordovician (Darriwilian)

Winneshiek Lagerstätte

 United States

Charbelicaris[51]

Gen. et sp. nov

Valid

Haug et al.

Late Cretaceous (Cenomanian)

Hadjoula Lagerstätte

 Lebanon

A member of Achelata. The type species is Charbelicaris maronites.

Corallioplax[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Goneplacoidea and the family Euryplacidae. The type species is C. exigua.

Costacopluma mamethioupamei[52]

Sp. nov

Valid

Hyžný et al.

Late Cretaceous (Maastrichtian)

Cap de Naze Formation

 Senegal

A retroplumid crab, a species of Costacopluma.

Cracchidynomene[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Dynomenidae. The type species is C. areolata.

Dardanus bayani[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A species of Dardanus.

Dardanus colosseus[53]

Sp. nov

Valid

Fraaije & Polkowsky

Eocene

 Austria

A hermit crab, a species of Dardanus.

Dardanus suessi[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A species of Dardanus.

Discutiolira[49]

Gen. et comb. nov

Valid

Robins et al.

Late Jurassic

 Austria
 Czech Republic
 Poland
 Romania

A member of Galatheoidea. The type species is "Galathea" eutecta Moericke (1899); genus also includes "Galathea" moesica Muţiu & Bădăluţă (1971).

Disipia[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A porcelain crab. The type species is D. sorbinii.

Dromilites montenati[54]

Sp. nov

Valid[55]

Robin et al.

Paleocene (Danian)

 France

A member of Dromiidae.

Dromiopsis ceratoi[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Dromiidae.

Dromiopsis longitudovata[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Dromiidae.

Dromiopsis parvula[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Dromiidae.

Dromiopsis vicetinus[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab.

Dynomene vetusta[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Dynomenidae.

Elektrocarcinus[36]

Gen. et comb. nov

Valid

Schweitzer et al.

Cretaceous (Albian to Maastrichtian)

 United Kingdom
 United States

A crab belonging to the group Raninoida and the family Necrocarcinidae. A new genus for "Necrocarcinus" pierrensis Rathbun (1917); genus also includes Elektrocarcinus davisi (Bishop, 1985), E. olsonorum (Bishop and Williams, 1991), E. texensis (Rathbun, 1935) and E. woodwardi (Bell, 1863).

Enoploclytia augustobonae[56]

Sp. nov

Valid

Devillez et al.

Early Cretaceous (Barremian)

 France

A member of Erymidae.

Enoploclytia gigantea[56]

Sp. nov

Valid

Devillez et al.

Early Cretaceous (Albian)

 United States

A member of Erymidae.

Eoacanthacaris[44]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A lobster. Genus includes new species E. tethysianus.

Eocarpilius lessineus[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Carpiliidae.

Eocharybdis rugosa[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Portunidae.

Eogeryon[57]

Gen. et sp. nov

Valid

Ossó

Late Cretaceous (Cenomanian)

 Spain

A member of Portunoidea. The type species is E. elegius.

Eomunidopsis prealpina[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A squat lobster belonging to the family Munidopsidae.

Eosadayoshia[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A member of Munididae. The type species is E. bolcensis.

Eoxanthops[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Xanthidae. The type species is E. scutatus.

Eryma vocontii[56]

Sp. nov

Valid

Devillez et al.

Early Cretaceous (Albian)

 France

A member of Erymidae.

Eumorphactaea convexa[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Pilumnidae.

Eurotanais pyrenaensis[35]

Sp. nov

Valid

Sánchez-García et al.

Late Cretaceous (Cenomanian)

Pyrenean amber

 France

A member of Tanaidacea belonging the family Alavatanaidae.

Eurotanais seilacheri[35]

Sp. nov

Valid

Sánchez-García et al.

Late Cretaceous (Turonian)

Vendean amber

 France

A member of Tanaidacea belonging the family Alavatanaidae.

Faxegalathea valeccensis[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A squat lobster. Originally assigned to the family Munidopsidae and to the genus Faxegalathea; subsequently transferred to the genus Tethysmunida by de Angeli & Ceccon (2017),[58] but this transfer was rejected by Beschin et al. (2019).[59]

Folguerolesia[60]

Gen. et comb. nov

Valid

Artal & Hyžný

Middle Eocene

 Spain

A member of Leucosioidea belonging to the new family Folguerolesiidae; a new genus for "Typilobus" boscoi Vía Boada (1959).

Fornicaris[61]

Gen. et sp. nov

Valid

Wilson & Selden in Selden et al.

Late Triassic (Norian)

Dolomia di Forni Formation

 Italy

A member of Asellota belonging to the family Paramunnidae. The type species is Fornicaris calligarisi.

Frontelata[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Carpilioidea and the family Palaeoxanthopsidae. The type species is F. spinacomposita.

Gabaleryon[62]

Gen. et 2 sp. et comb. nov

Valid[63]

Audo et al.

Early Jurassic (Toarcian)

 France
 Germany
 Italy
 United Kingdom

A member of Polychelida. Genus includes new species Gabaleryon coquelae and Gabaleryon garassinoi, as well as "Eryon" moorei.

Galathea lovarica[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A species of Galathea.

Galatheites aiola[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones
Štramberk Limestones

 Austria
 Czech Republic

A member of Galatheoidea, a species of Galatheites.

Galatheites diasema[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones

 Austria

A member of Galatheoidea, a species of Galatheites.

Galatheites obtecta[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

 Romania

A member of Galatheoidea, a species of Galatheites.

Gecchelicarcinus zanderigoi[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Macropipidae.

Glabropilumnus trispinosus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Pilumnidae.

Glyphithyreus bendensis[34]

Sp. nov

Valid

Schweitzer, Odumodu & Feldmann

Eocene

 Nigeria

A crab.

Graptocarcinus eocenicus[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Dynomenidae.

Haydnella granosa[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Xanthidae.

Hispanigalathea tithonia[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones

 Austria

A member of Galatheoidea, a species of Hispanigalathea.

Hohensteiniella[64]

Gen. et sp. nov

Valid[65]

Haug et al.

Early Devonian

Hunsrück Slate

 Germany

A crustacean with eumalacostracan affinities. The type species is H. engeli.

Iphiculus eliasi[66]

Sp. nov

Valid

Hyžný & Gross

Miocene

Florian Beds

 Austria
 Portugal[67]
 Spain[67]

A crab belonging to the superfamily Leucosioidea and the family Iphiculidae.

Iraticaris[68]

Gen. et sp. nov

Valid

Adami-Rodrigues & Pazinato in Adami-Rodrigues, Pazinato & Pinto

Early Permian

Irati Formation

 Brazil

A member of Pygocephalomorpha. The type species is I. damianii.

Karyosia[33]

Gen. et sp. nov

Valid

Schweitzer et al.

Early Cretaceous (Albian)

Glen Rose Formation
Walnut Formation

 United States

A member of Etyiidae. The type species is Karyosia apicava.

Kromtitis pseudolothi[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab.

Laevicarcinus serratus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Panopeidae.

Lemacola[49]

Gen. et 3 sp. nov

Valid

Robins et al.

Late Jurassic

 Austria
 Romania

A member of Galatheoidea. The type species is Lemacola jenniferae; genus also includes L. rossi and L. salia

Lessinipagurus boschettensis[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A hermit crab.

Lessinipagurus vasjamikuzi[69]

Sp. nov

Valid

Gašparič et al.

Eocene (late Bartonian–early Priabonian)

 Croatia

A hermit crab related to members of the genus Xylopagurus.

Libinia amplissimus[70]

Sp. nov

Valid

Feldmann & Schweitzer

Miocene

St. Marys Formation

 United States

A species of Libinia.

Liocarcinus priscus[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Carcinidae.

Lovaracarpilius[44]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Carpiliidae. The type species is L. incisus.

Lovaroides[44]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the superfamily Goneplacoidea and the family Carinocarcinoididae. The type species is L. elegans.

Macrocheira columbiaensis[71]

Sp. nov

Valid

Nyborg et al.

Early to middle Miocene

Astoria Formation

 United States

A relative of the Japanese spider crab.

Macrocheira jayi[71]

Sp. nov

Valid

Nyborg et al.

Late Eocene to Oligocene

Hesquiat Formation

 Canada

A relative of the Japanese spider crab.

Macrocheira sullivani[71]

Sp. nov

Valid

Nyborg et al.

Late Eocene to early Oligocene

Keasey Formation

 United States

A relative of the Japanese spider crab.

Magyarcarcinus yebraensis[72]

Sp. nov

Valid

Domínguez & Ossó

Eocene (Bartonian)

Margas de Arguís Formation

 Spain

A crab belonging to the superfamily Goneplacoidea, assigned to the new family Magyarcarcinidae.

Mesogalathea macra[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones

 Austria

A member of Galatheoidea, a species of Mesogalathea.

Mesogalathea pyxis[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones

 Austria

A member of Galatheoidea, a species of Mesogalathea.

Mesogalathea retusa[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones

 Austria

A member of Galatheoidea, a species of Mesogalathea.

Metopocarcinus fragilis[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Panopeidae.

Miopipus zovensis[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Carcinidae.

Mithracia lovatoi[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Majidae.

Montemagralia[73]

Gen. et sp. nov

Valid

De Angeli & Ceccon

Eocene (Ypresian)

 Italy

A crab belonging to the family Trapeziidae. The type species is M. lata.

Nanocassiope secretanae[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Xanthidae, a species of Nanocassiope.

Neocallichirus khadroensis[74]

Sp. nov

Valid

Hyžný & Charbonnier in Hyžný et al.

Paleocene (Danian)

Khadro Formation

 Pakistan

A member of Callianassidae, a species of Neocallichirus.

Neocallichirus lakhraensis[74]

Sp. nov

Valid

Hyžný & Charbonnier in Hyžný et al.

Eocene (Ypresian)

Lakhra Formation

 Pakistan

A member of Callianassidae, a species of Neocallichirus.

Nicoliscarcinus[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Cancridae. The type species is N. rotundatus.

Ophthalmoplax andina[75]

Sp. nov

Valid

Guzmán et al.

Late Cretaceous (late Campanian)

Lodolitas de Aguacaliente Formation

 Colombia

A member of Macropipidae, a species of Ophthalmoplax.

Paguristes paucituberculatus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A species of Paguristes.

Palaeodromites xestos[33]

Sp. nov

Valid

Schweitzer et al.

Early Cretaceous (Albian)

Walnut Formation

 United States

A member of Homolodromioidea belonging to the family Goniodromitidae, a species of Palaeodromites.

Palmyria levigata[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Domeciidae.

Panopeus postalensis[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Panopeidae, a species of Panopeus.

Paracancrinos[51]

Gen. et comb. nov

Valid

Haug et al.

Late Cretaceous (Cenomanian)

 Lebanon

A member of Achelata. A new genus for "Cancrinos" libanensis Garassino & Schweigert (2006).

Paracapsulapagurus[76]

Gen. et sp. nov

Valid

Hyžný et al.

Late Cretaceous (Maastrichtian)

Cape de Naze Formation

 Senegal

A hermit crab. The type species is P. poponguinensis.

Paracorallomursia[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Calappidae. The type species is P. medizzai.

Paradistefania denticulata[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Homolodromioidea and the family Goniodromitidae.

Paradynomene antiqua[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Dynomenidae.

Paragalathea arcella[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones

 Austria

A member of Galatheoidea, a species of Paragalathea.

Paragalathea crenarvina[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones
Štramberk Limestones

 Austria
 Czech Republic

A member of Galatheoidea, a species of Paragalathea.

Paragalathea ternata[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones

 Austria

A member of Galatheoidea, a species of Paragalathea.

Paragalathea vultuosona[49]

Sp. nov

Valid

Robins et al.

Late Jurassic

Ernstbrunn Limestones

 Austria

A member of Galatheoidea, a species of Paragalathea.

Paraocalina silviae[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Carpiliidae.

Paraporcellana[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A porcelain crab. The type species is P. fabianii.

Parhalimede[44]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Galenidae. The type species is P. ornata.

Parthenope menini[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Parthenopidae.

Permocaris[68]

Gen. et sp. nov

Valid

Adami-Rodrigues, Pazinato & Pinto

Early Permian

Irati Formation

 Brazil

A member of Pygocephalomorpha. The type species is P. purperae.

Petrochirus minutus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A member of Diogenidae.

Petrolisthes lineatus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A porcelain crab, a species of Petrolisthes.

Petrusia[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Plagusiidae. The type species is P. striata.

Phlyctenodes edwardsi[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Xanthidae, a species of Phlyctenodes.

Pilummede[77]

Gen. et sp. et comb. nov

Valid

Ossó & Clements

Eocene

Castle Hayne Limestone Formation
Clairbone Group

 United States

A member of Pilumnoidea of uncertain phylogenetic placement. The type species is P. penderensis; genus also includes "Lobonotus" bakeri Rathbun (1935).

Pilumnomimus dorsocarinatus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Xanthoidea of uncertain phylogenetic placement.

Pilumnomimus miettoi[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Xanthoidea of uncertain phylogenetic placement.

Pittinucaris[68]

Gen. et sp. nov

Valid

Adami-Rodrigues, Pazinato & Pinto

Early Permian

Irati Formation

 Brazil

A member of Pygocephalomorpha. The type species is P. wuerdigae.

Planocorystes[78]

Gen. et sp. nov

Valid

Franţescu, Feldmann & Schweitzer

Cretaceous

 United States

A crab belonging to the group Raninoida. The type species is P. robreidi.

Prealpicarcinus laisensis[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Pilumnidae.

Prealpiplax[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the superfamily Goneplacoidea and the family Euryplacidae. The type species is P. lessinea.

Prolecythocaris[79]

Gen. et sp. nov

Valid

Schweigert & Robins

Late Jurassic (early Kimmeridgian)

 Germany

A crab belonging to the family Lecythocaridae. The type species is P. hauckei.

Protomunida pentaspinosa[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A member of Munididae.

Pseudocorallomursia[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Calappidae. The type species is P. barbierii.

Pseudolambrus antiquus[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Parthenopidae.

Pustulina colossea[56]

Sp. nov

Valid

Devillez et al.

Early Cretaceous (Hauterivian)

 France

A member of Erymidae.

Pustulina occitana[56]

Sp. nov

Valid

Devillez et al.

Early Cretaceous (Berriasian)

 France

A member of Erymidae.

Rama[32]

Gen. et sp. nov

Junior homonym

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Cancridae. The type species is R. lineatuberculata. The generic name is preoccupied by Rama Bleeker; de Angeli & Ceccon (2017) coined a replacement name Ramacarcinus.[58]

Ramozius[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Pseudoziidae. The type species is R. punctatus.

Raniliformis clampensis[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Raninidae.

Rathbunopon viai[80]

Sp. nov

Valid

González-León et al.

Early Cretaceous (Albian)

 Spain

A homolodromioid crab belonging to the family Prosopidae.

Rosadromites[33]

Gen. et sp. nov

Valid

Schweitzer et al.

Early Cretaceous (Albian)

Glen Rose Formation

 United States

A member of Homolodromioidea belonging to the family Longodromitidae. The type species is Rosadromites texensis.

Rugosanilia[44]

Gen. et comb. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the family Raninidae. The type species is "Raniliformis" rugosa De Angeli & Beschin (2007).

Sakaila italica[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A species of Sakaila.

Santeexanthus caporiondoi[44]

Sp. nov

Valid

Beschin et al.

Eocene (Lutetian)

 Italy

A crab belonging to the superfamily Pseudozioidea and the family Pseudoziidae.

Scutata[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Tetraliidae. The type species is S. eocenica.

Serraphylctaena[49]

Gen. et comb. nov

Valid

Robins et al.

Cretaceous (Albian/Cenomanian)

 Spain

A member of Galatheoidea. The type species is "Paragalathea" multisquamata Vía Boada (1981).

Spathanomus[48]

Gen. et sp. nov

Valid

De Angeli

Eocene (Priabonian)

 Italy

A crab belonging to the group Cyclodorippoida and the family Cymonomidae. The type species is Spathanomus felicianensis.

Speocarcinus latus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Xanthidae, a species of Speocarcinus.

Tanidromites longinosa[81]

Sp. nov

Valid

Starzyk

Late Jurassic (Oxfordian)

 Poland

A homolodromioid crab belonging to the family Tanidromitidae.

Tanidromites muelleri[82]

Sp. nov

Valid

Krobicki & Zatoń

Middle Jurassic (late Bajocian)

 Poland

A homolodromioid crab belonging to the family Tanidromitidae.

Tanidromites schweitzerae[81]

Sp. nov

Valid

Starzyk

Late Jurassic (Oxfordian)

 Poland

A homolodromioid crab belonging to the family Tanidromitidae.

Tanidromites wysokaensis[81]

Sp. nov

Disputed

Starzyk

Late Jurassic (Oxfordian)

 Poland

A homolodromioid crab belonging to the family Tanidromitidae. Klompmaker et al. (2020) considered this species to be a junior synonym of Tanidromites scheffnerae Schweigert & Koppka (2011).[83]

Tethysastacus[56]

Gen. et comb. nov

Valid

Devillez et al.

Early Cretaceous (Valanginian)

 France

A member of Erymidae. The type species is T. tithonius.

Tethyscarpilius[84]

Gen. et sp. et comb. nov

Valid

De Angeli & Alberti

Eocene

 Italy
 United States

A crab belonging to the family Carpiliidae. Genus includes new species T. bericus, as well as "Palaeocarpilius" brodkorbi Lewis & Ross (1965).

Tetralia minuta[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Tetraliidae.

Texicancer[78]

Gen. et comb. nov

Valid

Franţescu, Feldmann & Schweitzer

Cretaceous

 United States

A crab belonging to the group Raninoida. The type species is "Necrocarcinus" renfroae Stenzel (1945).

Thalassina tsuyamensis[85]

Sp. nov

Valid

Ando & Kishimoto in Ando, Kishimoto & Kawano

Miocene

Yoshino Formation

 Japan

A species of Thalassina.

Thalassina yamato[85]

Sp. nov

Valid

Ando & Kishimoto in Ando, Kishimoto & Kawano

Miocene

Kawachi Formation

 Japan

A species of Thalassina.

Titanocarcinus schweitzerae[54]

Sp. nov

Valid[55]

Robin et al.

Paleocene (Danian)

 France

A crab belonging to the family Tumidocarcinidae.

Tonneleryon[86]

Gen. et sp. nov

Valid

Audo

Early Jurassic (Toarcian)

Posidonia Shale

 Germany

A member of Polychelida. The type species is T. schweigerti.

Tropicalia[32]

Gen. et sp. nov

Junior homonym

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Domeciidae. The type species is T. parva. The generic name is preoccupied by Tropicalia Kocak & Kemal (2008); de Angeli & Ceccon (2017) coined a replacement name Proticalia.[58]

Tuberosagalathea[49]

Gen. et comb. et 2 sp. nov

Valid

Robins et al.

Late Jurassic and Early Cretaceous (Hauterivian)

Ernstbrunn Limestones
Štramberk Limestones

 Austria
 Czech Republic
 France
 Poland
 Romania

A member of Galatheoidea. The type species is "Galathea" neojurensis Patrulius (1959); genus also includes "Galatheites" neocomiensis Van Straelen (1936) and "Eomunidopsis" portlandica Fraaye & Collins (1996), as well as new species Tuberosagalathea tornatilis and Tuberosagalathea antefixa.

Tymolus italicus[32]

Sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the group Cyclodorippoida and the family Cyclodorippidae.

Tytthotanais[35]

Gen. et sp. nov

Valid

Sánchez-García et al.

Late Cretaceous (Cenomanian)

Pyrenean amber

 France

A member of Tanaidacea of uncertain phylogenetic placement. The type species is T. tenvis.

Vasconilia[49]

Gen. et comb. et sp. nov

Valid

Robins et al.

Late Jurassic and Early Cretaceous

Ernstbrunn Limestones
Štramberk Limestones

 Austria
 Spain
 Japan?

A member of Galatheoidea. The type species is "Galathea" ruizi Van Straelen (1940); genus also includes "Galatheites" straeleni Ruiz de Gaona (1943), as well as new species Vasconilia xystos. Genus might also include ?Vasconilia miyakoensis (Takeda & Fujiyama, 1983).

Ypresicorystes[32]

Gen. et sp. nov

Valid

Beschin et al.

Eocene (Ypresian)

 Italy

A crab belonging to the family Corystidae. The type species is Y. expansus.

Zygastrocarcinus tricki[87]

Sp. nov

Valid

Nyborg et al.

Late Cretaceous (Campanian)

Pierre Shale

 United States

A crab belonging to the family Homolidae.

Ostracods

Name Novelty Status Authors Age Unit Location Notes Images

Actinocythereis imbeensis[88]

Sp. nov

Valid

De Mattos Manica & Coimbra

Early Miocene

Pelotas Basin

 Brazil

A member of the family Trachyleberididae, a species of Actinocythereis.

Amphizona? argentinensis[89]

Sp. nov

Valid

Salas

Devonian (Lochkovian)

Talacasto Formation

 Argentina

A member of Podocopa belonging to the Beyrichiocopida and the family Arcyzonidae.

Aparchitellina magnicornigera[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Beyrichicopida belonging to the family Aparchitellinidae.

Aparchitellina triangulata[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Rusanov Formation

 Russia

A member of Beyrichicopida belonging to the family Aparchitellinidae.

Asciocythere raybatei[91]

Nom. nov

Valid

Kempf

Middle Jurassic

 United Kingdom

A member of Schulerideidae; a replacement name for Asciocythere acuminata Bate (1964).

Bairdia sudrensis[92]

Sp. nov

Valid

Ied & Ismail

Miocene

 Egypt

A member of the family Bairdiidae.

Bingeria fluida[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Krasnaya Bukhta Formation
Ust’-Spokoinaya Formation

 Russia

A member of Beyrichicopida belonging to the family Beyrichiidae.

Bingeria lucida[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Srednii Formation

 Russia

A member of Beyrichicopida belonging to the family Beyrichiidae.

Bollia talacastensis[89]

Sp. nov

Valid

Salas

Devonian (Lochkovian)

Talacasto Formation

 Argentina

A member of Podocopa belonging to the Beyrichiocopida and the family Bollidae.

Brachycythere smithsoniana[93]

Sp. nov

Valid

Antonietto et al.

Early Cretaceous (Aptian-Albian)

Riachuelo Formation

 Brazil

A member of Podocopida belonging to the group Cytherocopina and the family Trachyleberididae.

Buntonia bassiounii[94]

Sp. nov

Valid

Morsi, Hewaidy & Samir

Eocene

 Egypt

A member of Trachyleberididae, a species of Buntonia.

Buntonia posteroacuta[94]

Sp. nov

Valid

Morsi, Hewaidy & Samir

Eocene

 Egypt

A member of Trachyleberididae, a species of Buntonia.

Bythoceratina brunomilhaui[91]

Nom. nov

Valid

Kempf

Miocene

Waitemata Basin

 New Zealand

A member of Bythocytheridae; a replacement name for Bythoceratina robusta Milhau (1993).

Bythocythere agostinae[95]

Sp. nov

Valid

Sciuto

Pleistocene

 Italy

Candona csikiiformis[96]

Sp. nov

Valid

Teterina

Miocene

 Russia

Conchoprimitia brasiliensis[97]

Sp. nov

Valid

Adôrno & Salas in Adôrno et al.

Late Ordovician

Vila Maria Formation

 Brazil

A member of Podocopa belonging to the order Beyrichiocopida and the family Conchoprimitiidae.

Cryptophyllus fossularis[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Ust’-Spokoinaya Formation

 Russia

Cryptophyllus magnei[98]

Sp. nov

Valid

Maillet & Casier in Maillet, Danelian & Casier

Devonian (Givetian)

Fromelennes Formation

 Belgium

A possible member of Palaeocopida belonging to the family Rhabdostichidae, a species of Cryptophyllus.

Ctenoloculinella papillifera[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation
Rusanov Formation

 Russia

A member of Hollinocopida belonging to the family Ctenoloculinidae.

Cytherella abdelshafyii[92]

Sp. nov

Valid

Ied & Ismail

Miocene

 Egypt

A member of the family Cytherellidae.

Cytherellina subelliptica[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation
Rusanov Formation

 Russia

A member of Metacopida belonging to the family Healdiidae.

Cytheretta africana[92]

Sp. nov

Valid

Ied & Ismail

Miocene

 Egypt

A member of the family Cytherettidae.

Cytheretta sinaensis[92]

Sp. nov

Valid

Ied & Ismail

Miocene

 Egypt

A member of the family Cytherettidae.

Cytheropteron morsii[91]

Nom. nov

Valid

Kempf

Eocene

 Egypt

A member of Cytheruridae, a species of Cytheropteron. The original specific name was Cytheropteron bicostatum, Morsi, Hewaidy & Samir (2016),[94] which turned out to be preoccupied by Cytheropteron bicostatum Brand (1990).

Cytheropteron nukhulensis[94]

Sp. nov

Valid

Morsi, Hewaidy & Samir

Eocene

 Egypt

A member of Cytheruridae, a species of Cytheropteron.

Cytheropteron speijeri[94]

Sp. nov

Valid

Morsi, Hewaidy & Samir

Eocene

 Egypt

A member of Cytheruridae, a species of Cytheropteron.

Digmocythere centroreticulata[94]

Sp. nov

Valid

Morsi, Hewaidy & Samir

Eocene

 Egypt

A member of Trachyleberididae, a species of Digmocythere.

Ectonodoconcha[99]

Gen. et sp. nov

Valid

Ceolin & Whatley in Ceolin et al.

Paleocene (Danian)

Neuquén Basin

 Argentina

A member of Cytheridae. The type species is Ectonodoconcha lepidotus.

Eobeyrichia nordensis[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Srednii Formation

 Russia

A member of Beyrichicopida belonging to the family Beyrichiidae.

Eobeyrichia subeurina[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Samoilovich Formation
Srednii Formation

 Russia

A member of Beyrichicopida belonging to the family Beyrichiidae.

Eokloedenia reticulata[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Late Silurian

Krasnaya Bukhta Formation

 Russia

A member of Beyrichicopida belonging to the family Kloedeniidae.

Eukloedeneterra[90]

Gen. et sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Samoilovich Formation
Srednii Formation

 Russia

A member of Kloedenellocopida belonging to the family Kloedenellidae. The type species is E. borealis.

Evlanella peranensis[100]

Sp. nov

Valid

Maillet in Maillet et al.

Devonian (Givetian)

Candás Formation

 Spain

A species of Evlanella.

Evlanella tuberculifera[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Early Devonian

Al’banov Formation

 Russia

A member of Kloedenellocopida belonging to the family Lichviniidae.

Gabonorygma[93]

Gen. et comb. et sp. nov

Valid

Antonietto et al.

Late Jurassic (Kimmeridgian–early Tithonian) and Early Cretaceous (Aptian-Albian)

Madiéla Formation
Riachuelo Formation

 Brazil
 Denmark
 Gabon

A member of Podocopida belonging to the group Cytherocopina and the family Krithidae. The type species is "Dicrorygma" brotzeni Christensen (1965); genus also includes new species G. sergipana.

Kozlowskiella? nuda[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Beyrichicopida belonging to the family Aparchitellinidae.

Kuznetsoviella[101][102]

Nom. nov

Valid

Doweld

Early Cretaceous (Barremian)

 Azerbaijan

A replacement name for Aenigma Kuznetsova (1957) and Kuznetsovia Doweld (2016) (both preoccupied).

Limnocythere alexanderi[103]

Sp. nov

Valid

Palacios-Fest, Cusminsky & McGlue

Late Quaternary

 Argentina

A species of Limnocythere.

Limnocythere foresteri[103]

Sp. nov

Valid

Palacios-Fest, Cusminsky & McGlue

Late Quaternary

 Argentina

A species of Limnocythere.

Limnocythere lysandrosi[103]

Sp. nov

Valid

Palacios-Fest, Cusminsky & McGlue

Late Quaternary

 Argentina

A species of Limnocythere.

Limnocythere ruipunctifinalis[103]

Sp. nov

Valid

Palacios-Fest, Cusminsky & McGlue

Late Quaternary

 Argentina

A species of Limnocythere.

Microxestoleberis scillae[95]

Sp. nov

Valid

Sciuto

Pleistocene

 Italy

Mongolocypris kohi[104]

Sp. nov

Valid

Choi & Huh

Early Cretaceous (Albian)

Jinju Formation

 South Korea

A non-marine ostracod, a species of Mongolocypris.

Mosolovina conifera[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Beyrichicopida belonging to the family Aparchitellinidae.

Mosolovina costulifera[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Beyrichicopida belonging to the family Aparchitellinidae.

Neohornibrookella nepeani[105]

Sp. nov

Valid

Warne & Whatley

Neogene

 Australia

Neurocythere multiforma[106]

Sp. nov

Valid

Pais et al.

Middle Jurassic (Callovian)

 Portugal

Nodoconcha polytorosa[99]

Sp. nov

Valid

Ceolin & Whatley in Ceolin et al.

Paleocene (Danian)

Neuquén Basin

 Argentina

A member of Cytheridae, a species of Nodoconcha.

Nodoconcha sanniosis[99]

Sp. nov

Valid

Ceolin & Whatley in Ceolin et al.

Paleocene (Danian)

Neuquén Basin

 Argentina

A member of Cytheridae, a species of Nodoconcha.

Nodoconcha upsilon[99]

Sp. nov

Valid

Ceolin & Whatley in Ceolin et al.

Paleocene (Danian)

Neuquén Basin

 Argentina

A member of Cytheridae, a species of Nodoconcha.

Norditerria[90]

Gen. et 2 sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Krasnaya Bukhta Formation
Ust’-Spokoinaya Formation

 Russia

A member of Beyrichicopida belonging to the family Hexophthalmoididae. The type species is N. torosa; genus also includes N. simplex.

Noviportia cornuta[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Samoilovich Formation
Srednii Formation

 Russia

A member of Beyrichicopida belonging to the family Zygobolbidae.

Nyhamnella elongata[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Srednii Formation

 Russia

A member of Kloedenellocopida belonging to the family Gotlandellidae.

Occultocythereis alsheikhlyi[91]

Nom. nov

Valid

Kempf

Late Cretaceous (Maastrichtian)

 Iraq

A member of Trachyleberididae; a replacement name for Occultocythereis elongata Al-Sheikhly (1982).

Oertliella petraensis[107]

Sp. nov

Valid

Al-Sheikhly

Paleocene (Danian)

Taqiye Formation

 Jordan

A member of the family Trachyleberididae, a species of Oertliella.

Paenaequina recta[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Devonian (Emsian)

Al’banov Formation

 Russia

A member of Leperditicopida belonging to the family Schrenckiidae.

Palaeocytheridea (Malzevia)? dorsocostata[106]

Sp. nov

Valid

Pais et al.

Middle Jurassic (Callovian)

 Portugal

Paragrenocythere monilis[107]

Sp. nov

Valid

Al-Sheikhly

Late Cretaceous (Maastrichtian)

Shiranish Formation

 Iraq

A member of the family Trachyleberididae, a species of Paragrenocythere.

Parakrithe tayibaensis[94]

Sp. nov

Valid

Morsi, Hewaidy & Samir

Eocene

 Egypt

A member of Krithidae, a species of Parakrithe.

Peloriops levisulcata[107]

Sp. nov

Valid

Al-Sheikhly

Late Cretaceous (late Campanian?-Maastrichtian)

Hartha Formation
Shiranish Formation

 Iraq

A member of the family Trachyleberididae, a species of Peloriops.

Perissocytheridea colini[108]

Sp. nov

Valid

Nogueira & Ramos

Oligocene-Miocene

Pirabas Formation

 Brazil

An ostracod, a species of Perissocytheridea.

Perissocytheridea largulateralis[108]

Sp. nov

Valid

Nogueira & Ramos

Oligocene-Miocene

Pirabas Formation

 Brazil

An ostracod, a species of Perissocytheridea.

Perissocytheridea pirabensis[108]

Sp. nov

Valid

Nogueira & Ramos

Oligocene-Miocene

Pirabas Formation

 Brazil

An ostracod, a species of Perissocytheridea.

Perissocytheridea punctoreticulata[108]

Sp. nov

Valid

Nogueira & Ramos

Oligocene-Miocene

Pirabas Formation

 Brazil

An ostracod, a species of Perissocytheridea.

Petrisigmoopsis? rotundum[89]

Sp. nov

Valid

Salas

Devonian (Lochkovian)

Los Espejos Formation
Talacasto Formation

 Argentina

A member of Podocopa belonging to the Beyrichiocopida and the family Bollidae.

Pircawayra antiqua[89]

Sp. nov

Valid

Salas

Devonian (Lochkovian)

Talacasto Formation

 Argentina

A member of Podocopa belonging to the Beyrichiocopida and the superfamily Kloedenelloidea.

Platybolbina calva[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Late Silurian

Krasnaya Bukhta Formation

 Russia

A member of Hollinocopida belonging to the family Eurychilinidae.

Praebythoceratina deltalata[93]

Sp. nov

Valid

Antonietto et al.

Early Cretaceous (Aptian-Albian)

Riachuelo Formation

 Brazil

A member of Podocopida belonging to the group Cytherocopina and the family Bythocytheridae.

Praebythoceratina parascrobiculata[106]

Sp. nov

Valid

Pais et al.

Middle Jurassic (Callovian)

 Portugal

Praeschuleridea lisensis[106]

Sp. nov

Valid

Pais et al.

Middle Jurassic (Callovian)

 Portugal

Protobrachycythere[109]

Gen. et 2 sp. nov

Valid

Karpuk

Early Cretaceous (Barremian-Aptian)

Crimean Peninsula

A member of the family Protocytheridae. The type species is P. taurica; genus also includes P. aptica.

Pteriella? dubia[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Srednii Formation

 Russia

A member of Hollinocopida belonging to the suborder Primitiopsicopina.

Reticulina ninurta[107]

Sp. nov

Valid

Al-Sheikhly

Middle-late Eocene

Avanah Formation
Jaddala Formation

 Iraq

A member of the family Trachyleberididae, a species of Reticulina.

Reticulina syriaensis[107]

Sp. nov

Valid

Al-Sheikhly

Paleocene

Aaliji Formation

 Syria

A member of the family Trachyleberididae, a species of Reticulina.

Saccarchites tumefactus[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Late Silurian

Ust’-Spokoinaya Formation

 Russia

A member of Beyrichicopida belonging to the family Ochescaphidae.

Samarella centrigibbera[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Rusanov Formation

 Russia

A member of Metacopida belonging to the family Rishonidae.

Sapucariella[110]

Gen. et comb. et 3 sp. nov

Valid

Puckett, Andreu & Colin

Early Cretaceous (Aptian)-early Paleocene

Agulhas Bank
 Algeria
 Brazil
 Cameroon
 Egypt
 Gabon
 Ghana
 India
 Iran
 Israel
 Ivory Coast
 Kuwait
 Lebanon
 Libya
 Madagascar
 Mali
 Morocco
 Niger
 Nigeria
 Oman
 Saudi Arabia
 Senegal
 South Africa
 Tanzania
 Togo
 Tunisia
 United Arab Emirates

A member of the family Brachycytheridae. The type species is "Brachycythere" sapucariensis Krömmelbein (1964); genus also includes "Brachycythere" agulhasensis Dingle (1971), S. angulata (Grékoff, 1951), "Brachycythere" arabica Al-Furaih (1985), "Brachycythere" armata Reyment (1960), "Brachycythere" beershevaensis Honigstein (1984), "Brachycythere" dumoni Bismuth & Saint-Marc in Bismuth et al. (1981), "Brachycythere" ekpo Reyment (1960), "Brachycythere" glypta Al-Furaih (1985), "Brachycythere" ilamensis Emami (1990), "Brachycythere" iranensis Emami (1990), "Brachycythere" jodhpurensis Singh (1997), "Brachycythere" kulatturensis Guha (1971), "Brachycythere" labioforma Emami (1990), "Cytheridea" longicaudata Chapman (1904), "Brachycythere" meata Al-Furaih (1985), "Brachycythere" multidifferentis Nicolaidis & Piovesan in Piovesan et al. (2013), "Brachycythere" oguni Reyment (1960), "Brachycythere" posterotruncata Emami (1990), "Brachycythere" pucketti Babinot et al. (2009), "Brachycythere" reymenti Emami (1990), "Brachycythere" rotunda Dingle (1969), "Brachycythere" shamlani Al-Furaih (1985), "Brachycythere" sicarius Dingle (1980), "Brachycythere" sillakkudiensis Sastry et al. (1972), "Brachycythere" subtriangulata Singh & Porwal (1989), "Brachycythere" tumida Al-Furaih (1985), "Brachycythere" undosa Al-Furaih (1985) and "Brachycythere" ventrocomplanatus Delicio et al. (2000), as well as new species S. honigsteini, S. parvoangulata and S. pseudosapucariensis.

Satiellina paranaensis[97]

Sp. nov

Valid

Adôrno & Salas in Adôrno et al.

Late Ordovician

Vila Maria Formation

 Brazil

A member of Podocopa belonging to the order Beyrichiocopida and the superfamily Drepanelloidea.

Sibiritia ? septentrionalis[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Golomyannyi Formation
Srednii Formation
Vodopad Formation

 Russia

A member of Leperditicopida belonging to the family Sibiritiidae.

Signetopsis bolbilobata[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Ust’-Spokoinaya Formation

 Russia

A member of Hollinocopida belonging to the family Signetopsidae.

Suinella[89]

Gen. et sp. nov

Valid

Salas

Devonian (Lochkovian)

Talacasto Formation

 Argentina

A member of Podocopa belonging to the Beyrichiocopida and possibly to the family Primitiopsidae. The type species is S. huarpesi.

Sulcella? acuminata[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Kloedenellocopida belonging to the family Mennerellidae.

Sulcella? annae[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Kloedenellocopida belonging to the family Mennerellidae.

Sulcella? modesta[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Kloedenellocopida belonging to the family Mennerellidae.

Timanella magna[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Platycopida belonging to the family Ellesclavidae.

Timanella rara[90]

Sp. nov

Valid

Evdokimova in Abushik & Evdokimova

Devonian

Al’banov Formation

 Russia

A member of Platycopida belonging to the family Ellesclavidae.

Tollitia navicula[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Golomyannyi Formation
Srednii Formation

 Russia

A member of Leperditicopida belonging to the family Tollitiidae.

Tricostabrachycythere[110]

Gen. et comb. nov

Valid

Puckett, Andreu & Colin

Cretaceous (Albian-Cenomanian)

Goru Formation

 India
 Morocco
 Venezuela

A member of the family Brachycytheridae. The type species is "Veenia" kharatarensis Singh (1997).

Tvaerenella? ambigua[90]

Sp. nov

Valid

Abushik in Abushik & Evdokimova

Silurian

Srednii Formation

 Russia

A member of Hollinocopida belonging to the family Piretellidae.

Virgulacytheridea posteroacuminata[106]

Sp. nov

Valid

Pais et al.

Middle Jurassic (Callovian)

 Portugal

Xestoleberis morsiana[91]

Nom. nov

Valid

Kempf

Eocene

 Egypt

A member of Xestoleberididae, a species of Xestoleberis. The original specific name was Xestoleberis posterotruncata Morsi, Hewaidy & Samir (2016),[94] which turned to be preoccupied by Xestoleberis posterotruncata Titterton & Whatley (2005).

Other crustaceans

Name Novelty Status Authors Age Unit Location Notes Images

Apocyclops californicus[111]

Sp. nov

Valid

Hołyńska, Leggitt & Kotov

Miocene (latest Burdigalian or earliest Langhian)

Barstow Formation

 United States

A cyclopid copepod, a species of Apocyclops.

Arcuatoscalpellum[112]

Gen. et comb. nov

Valid

Gale

Cretaceous (Aptian to Maastrichtian)

 France
 United Kingdom

A goose barnacle belonging to the family Scalpellidae. The type species is "Scalpellum" arcuatum Darwin (1851); genus also includes "Scalpellum" (Arcoscalpellum) comptum Withers (1910) and "Scalpellum" trilineatum Darwin (1851).

Catherinum anglicum [112]

Sp. nov

Valid

Gale

Late Cretaceous (Campanian)

 United Kingdom

A goose barnacle belonging to the family Scalpellidae, a species of Catherinum.

Cetopirus fragilis[113]

Sp. nov

Valid

Collareta et al.

Early Pleistocene

 Italy

A whale barnacle, a species of Cetopirus.

Conopea bacata[114]

Sp. nov

Valid

Carriol in Carriol & Schneider

Miocene (Burdigalian)

 Germany

A barnacle belonging to the family Archaeobalanidae, a species of Conopea.

Gesvesia[115]

Gen. et sp. nov

Valid

Gueriau et al.

Devonian (Famennian)

 Belgium

A clam shrimp belonging to the group Spinicaudata. The type species is Gesvesia pernegrei.

Haltinnaias[115]

Gen. et sp. nov

Valid

Gueriau et al.

Devonian (Famennian)

 Belgium

A member of Anostraca. The type species is Haltinnaias serrata.

Myolepas reussi[116]

Sp. nov

Valid

Kočí et al.

Late Cretaceous (late Cenomanian to early Turonian)

Bohemian Cretaceous Basin

 Czech Republic

A goose barnacle.

Nestoria sikeshuensis[117]

Sp. nov

Valid

Teng et al.

Early Cretaceous

Tugulu Group

 China

A clam shrimp belonging to the superfamily Eosestherioidea and the family Nestoriidae.

Pachyscalpellum heltzeli[118]

Sp. nov

Valid

Jagt, Verhesen & Goolaerts

Late Cretaceous (Maastrichtian)

El Haria Formation

 Tunisia

A goose barnacle belonging to the family Scalpellidae, a species of Pachyscalpellum.

Plesiobrachylepas[119]

Gen. et sp. nov

Valid

Carriol in Carriol et al.

Eocene

Mo Clay Formation

 Denmark

A barnacle belonging to the group Sessilia and the family Brachylepadidae. The type species is Plesiobrachylepas jutlandica.

Pseudoscapholeberis[120]

Gen. et sp. nov

Valid

Flössner & Fryer

Cretaceous (late Albian or earliest Cenomanian)

 France

A member of Anomopoda. The type species is Pseudoscapholeberis enigmatica.

Scillaelepas danningeri[114]

Sp. nov

Valid

Carriol in Carriol & Schneider

Miocene (Burdigalian)

Ottnang Formation

 Austria
 Germany

A barnacle belonging to the family Calanticidae, a species of Scillaelepas.

Stipilepas[119]

Gen. et sp. nov

Valid

Carriol in Carriol et al.

Eocene

Mo Clay Formation

 Denmark

A goose barnacle belonging to the group Scalpelliformes and the family Eolepadidae. The type species is Stipilepas molerensis.

Insects

Trilobites

Research

  • Queues of the trilobite Trimerocephalus chopini are described from the Devonian (Famennian) of Poland by Błażejowski et al. (2016), who interpret these queues as representing mass migratory chains.[121]
  • Disarticulated exoskeletons of the phacopids belonging to the species Omegops cornelius preserved within a nautiloid conch are described from the Devonian Hongguleleng Formation (China) by Zong, Fan & Gong (2016), who interpret the fossils as indicating that the trilobites had moulted within the living chamber of a dead nautiloid.[122]

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Abarrandia[123]

Gen. et sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

A nileid. Genus includes new species A. wuhaiensis.

Acanthomicmacca franconica[124]

Sp. nov

Valid

Geyer

Cambrian

 Germany

A member of Chengkouiidae.

Acanthomicmacca hupei[124]

Sp. nov

Valid

Geyer

Middle Cambrian

Jbel Wawrmast Formation

 Morocco

A member of Chengkouiidae.

Acanthomicmacca mirandoides[124]

Sp. nov

Valid

Geyer

Cambrian

A member of Chengkouiidae.

Acanthomicmacca morrisoni[124]

Sp. nov

Valid

Geyer

Middle Cambrian

Jbel Wawrmast Formation

 Morocco

A member of Chengkouiidae.

Acanthomicmacca rana[124]

Sp. nov

Valid

Geyer

Cambrian

A member of Chengkouiidae.

Acastoides poschmanni[125]

Sp. nov

Valid

Basse & Müller

Devonian (Emsian)

Rupbach Shale

 Germany

Agasella prona[126]

Sp. nov

Valid

Salikhova in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Agraulidae.

Amphitryon xiushanensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Ampyx gongwusuensis[128]

Sp. nov

Valid

Lee et al.

Ordovician (Darriwilian)

Klimoli Formation

 China

Ampyxinella latilimbata[123]

Sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

Amslamia[124]

Gen. et sp. nov

Valid

Geyer

Cambrian

 Morocco

A member of Chengkouiidae. Genus includes new species A. rudkini.

Anaxarete[124]

Gen. et comb. nov

Junior homonym

Geyer

Cambrian

A member of Protoleninae; a new genus for "Micmacca" ellipsocephaloides var. senior (recombined as Anaxarete senior). The generic name is preoccupied by Anaxarete Gistl (1848).

Anechocephalus intermedius[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Anechocephalus rebeccaae[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Angustinouyia[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Inouyiidae.

Apatokephalus rugosus[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Asaphellus clarksoni[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Aulacopleura (Paraaulacopleura) lemkei[125]

Sp. nov

Valid

Basse & Müller

Devonian (Eifelian)

Rupbach Shale

 Germany

Austerops legrandi[132]

Sp. nov

Valid

Khaldi et al.

Early Devonian

‘Chefar el Ahmar’ Formation

 Algeria

A phacopid, a species of Austerops.

Australoscutellum talenti[133]

Sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

Barrandeops chattertoni[132]

Sp. nov

Valid

Khaldi et al.

Early Devonian

‘Chefar el Ahmar’ Formation

 Algeria

A phacopid. Originally described as a species of Barrandeops; Van Viersen & Holland (2016) considered the genus Barrandeops to be a junior synonym of the genus Morocops[134] and subsequently Van Viersen, Holland & Koppka (2017) transferred the species B. chattertoni to the genus Morocops.[135]

Barrandeops (Lahnops)[125]

Subgen. et sp. et nom. nov

Valid

Basse & Müller

Devonian

Leun Shale
Rupbach Shale

 Germany

The subgenus includes new species Barrandeops (Lahnops) steinmeyeri, as well as Barrandeops (Lahnops) burhennei (a replacement name for Phacops holzapfeli Burhenne).

Binodaspis nuda[126]

Sp. nov

Valid

Pegel in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Ptychopariidae.

Buitella concava[126]

Sp. nov

Valid

Pegel & Salikhova in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the subfamily Conokephalininae.

Camarella[130]

Gen. nov

Junior homonym

Zhu in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Solenopleuridae. The generic name is preoccupied by Camarella Dall (1877).

Carioides[136]

Gen. et sp. nov

Valid

Bushuev & Makarova

Cambrian

Chaya Formation

 Russia

A member of Anomocaridae. Genus includes new species C. enodis.

Cernuolimbus ludvigseni[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Chlupacops[137]

Gen. et sp. et comb. nov

Valid

Feist in Feist, Mahboubi & Girard

Late Devonian

 Algeria
 Germany
 Morocco

A member of Phacopidae. The type species is C. laticeps; genus also includes "Phacops" cryphoides Richter & Richter (1926) and possibly "Phacops" reichi Kegel (1931), "Phacops" koeneni Holzapfel (1895) and "Nephranops" spectabilis Meischner (1965).

Claycreekia[129]

Gen. et sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

A member of the family Olenidae. Genus includes new species C. chrisorum.

Cliffia aitkeni[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Corrugatagnostus ningqiangensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Cybantyx? ergodes[133]

Sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

Cyclopyge latilimbata[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Cyphaspides (Cyphaspides) malbertii[125]

Sp. nov

Valid

Basse & Müller

Devonian (Eifelian)

Rupbach Shale

 Germany

Cyphaspides (Cyphaspides) weugi[125]

Sp. nov

Valid

Basse & Müller

Devonian (Eifelian)

Wissenbach Shale

 Germany

Cyphaspis bluhmi[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Eifelian)

Taboumakhloûf Formation?

 Morocco

Cyphaspis boninoi[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Eifelian)

Hanonet Formation

 Belgium

Cyphaspis eximia[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Eifelian)

 Morocco

Cyphaspis foumzguidensis[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Eifelian)

Timrhanrhart Formation

 Morocco

Cyphaspis heisingi[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Pragian)

Assa Formation

 Morocco

Cyphaspis ihmadii[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Givetian)

Bou Dîb Formation

 Morocco

Cyphaspis juergenhollandi[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Emsian)

Amerboh Group

 Morocco

Cyphaspis khraidensis[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Emsian)

Timrhanrhart Formation

 Morocco

Cyphaspis kippingi[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Emsian)

Tazoulait Formation

 Morocco

Cyphaspis kweberi[125]

Sp. nov

Valid

Basse & Müller

Devonian (Emsian)

Rupbach Shale

 Germany

Cyphaspis lerougei[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Eifelian)

Khebchia Formation

 Morocco

Cyphaspis maharchensis[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Pragian)

Ihandar Formation

 Morocco

Cyphaspis smeenki[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Eifelian)

El Otfal Formation

 Morocco

Cyphaspis tadachachtensis[134]

Sp. nov

Valid

Van Viersen & Holland

Devonian (Emsian)

Khebchia Formation

 Morocco

Dechenella burmeisteri emstae[138]

Subsp. nov

Valid

Basse, Koch & Lemke

Devonian (Givetian)

 Germany

A member of the family Proetidae.

Degamella interrupta[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Degamella latifrons[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Destombesina schumacherorum[125]

Sp. nov

Valid

Basse & Müller

Devonian (Emsian)

Rupbach Shale

 Germany

Diademaproetus habenichti[125]

Sp. nov

Valid

Basse & Müller

Devonian (Emsian)

Rupbach Shale

 Germany

Dislobosaspis angustiglossoides[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Dolabrapex[133]

Gen. et sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

A member of Scutelluidae. Genus includes new species D. acomus.

Dorypyge? aliena[126]

Sp. nov

Valid

Pegel in Pegel et al.

Cambrian

 Russia

A member of Dorypygidae.

Dunderbergia? edgecombei[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Eccaparadoxides? thorslundi[139]

Sp. nov

Valid

Rushton, Weidner & Ebbestad

Cambrian

 Sweden

A member of Paradoxididae of uncertain phylogenetic placement, possibly a species of Eccaparadoxides.

Echidnops taphomimus[140]

Sp. nov

Valid

Rustán & Balseiro

Early Devonian

Talacasto Formation

 Argentina

Effnaspis clavata[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Eiluroides[130]

Gen. nov

Valid

Yuan & Zhu in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Menocephalidae.

Encrinurella latifrons[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Enigmapyge[137]

Gen. et sp. nov

Valid

Feist in Feist, Mahboubi & Girard

Devonian (early Famennian)

“Argiles de Marhouma” Formation

 Algeria

A member of Phacopidae. The type species is E. marhoumensis.

Erytheia[124]

Gen. et comb. nov

Valid

Geyer

Cambrian

 Poland

A member of Chengkouiidae. Genus includes "Micmacca" klimontowi Orłowski (1985).

Eugonocare? phillipi[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Forchhammeria insueta[126]

Sp. nov

Valid

Salikhova in Pegel et al.

Cambrian

 Russia

A member of Asaphida belonging to the family Anomocaridae.

Gaindella[126]

Gen. et sp. nov

Valid

Pegel in Pegel et al.

Cambrian

 Russia

A member of Asaphida belonging to the family Pterocephaliidae. The type species is G. angusta.

Galbagnostus obscurus[123]

Sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

Geesops fabrei[132]

Sp. nov

Valid

Khaldi et al.

Early Devonian

‘Chefar el Ahmar’ Formation

 Algeria

A phacopid. Originally described as a species of Geesops; Van Viersen, Holland & Koppka (2017) transferred the species to the genus Adrisiops.[135]

Gymnagnostus kobayashii[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Hadromeros hunanensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Helanshanaspis[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Ptychopariidae.

Hexacopyge scopulata[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Holdenia liangshanensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Hydrocephalus spinulosus[139]

Sp. nov

Valid

Rushton, Weidner & Ebbestad

Cambrian

 Sweden

A member of Paradoxididae.

Iguidiella[124]

Gen. et sp. nov

Valid

Geyer

Cambrian

A member of Chengkouiidae. Genus includes new species I. schaeri.

Illaenoscutellum psephos[133]

Sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

Illaenus slancyensis[141]

Sp. nov

Valid

Krylov

Ordovician

Kukruse Formation

 Russia

Inouyia (Sulcinouyia)[130]

Subgen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Inouyiidae.

Iotoryx[133]

Gen. et sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

A member of Scutelluidae. Genus includes new species I. clarksoni.

Irbuklina[126]

Gen. et 2 sp. nov

Valid

Egorova in Pegel et al.

Cambrian

 Russia

A member of Asaphida of uncertain phylogenetic placement. The type species is I. candida; genus also includes I. rigida.

Irvingella dawnae[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Irvingella lata[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Irvingella orrensis[142]

Sp. nov

Valid

Westrop & Adrain

Cambrian

Orr Formation

 United States

Japonoscutellum drakton[133]

Sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

Japonoscutellum fractum[133]

Sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

Japonoscutellum mawsonae[133]

Sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

Juliaspis convexa[126]

Sp. nov

Valid

Pegel in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Namanoiidae.

Kanlingia mirabilis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Kanlingia reshiensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Kanlingia rhombica[123]

Sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

Kendallina greenensis[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Kindbladia wilsoni[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Kodymaspis expansa[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Kodymaspis maocaopuensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Koneprusia martini[125]

Sp. nov

Valid

Basse & Müller

Devonian (Emsian)

Rupbach Shale

 Germany

Kosovopeltis avita[133]

Sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

Lamproscutellum scoparium[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Leiolonchodomas[127]

Gen. et sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Genus includes new species L. dangmengouensis.

Lenacare sentum[126]

Sp. nov

Valid

Salikhova in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Alokistocaridae.

Linguagnostus aldanicus[143]

Sp. nov

Valid

Makarova & Bushuev

Cambrian

Chaya Formation

 Russia

A member of Diplagnostidae.

Lioaspis[126]

Gen. et sp. nov

Valid

Lazarenko & Salikhova in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Asaphiscidae. The type species is L. infida.

Liopeishaniella[144]

Gen. et comb. nov

Valid

Bentley, Jago & Cooper

Cambrian (Drumian)

Antarctica
(Victoria Land)

A polymerid trilobite. Genus includes "Liopeishania" angusta Wolfart (1994).

Liopelta paula[126]

Sp. nov

Valid

Salikhova in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Anomocarellidae.

Liquanella[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Asaphiscidae.

Lisogorites yaoi[123]

Sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

An asaphid.

Lonchocheirurus[127]

Gen. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Lonchodomas yichongiaoensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Metayuepingia riccardii[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Microparia (Heterocyclopyge) obsoleta[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Microparia (Heterocyclopyge) taoyuanensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Mioptychopyge semicircula[123]

Sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

An asaphid.

Nileus cingolanii[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Nileus erici[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Niuchangella angustilimbata[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Niuchangella yuanbaensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Niuxinshania[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Crepicephalidae.

Oedicybele caeca[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Ogygiocaris? iruyensis[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Olenaspella chrisnewi[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Olenoides sassikaspa[144]

Sp. nov

Valid

Bentley, Jago & Cooper

Cambrian (Drumian)

Antarctica
(Victoria Land)

Onychopyge acenolazai[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Onychopyge gonzalezae[131]

Sp. nov

Valid

Tortello & Esteban

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

Opsypharus pandanensis[133]

Sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

Orientanomocare[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Anomocarellidae.

Orygmaspis (Parabolinoides) doriansmithi[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Orygmaspis (Parabolinoides) jenkinsi[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Orygmaspis (Parabolinoides) mckellari[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Orygmaspis (Parabolinoides) medispina[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Orygmaspis (Parabolinoides) morrisi[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Otarion fugitivum[145]

Sp. nov

Valid

McNamara & Feist

Devonian (Frasnian)

Virgin Hills Formation

 Australia

A proetid, a species of Otarion.

Ovalocephalus pyriformis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Ovalocephalus shaanxiensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Palaeosunaspis[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Sunaspidae.

Panderia acutata[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Panderia lubrica[123]

Sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

Parabolinella clarisae[146]

Sp. nov

Valid

Monti, del Huerto Benítez & Ramirez

Ordovician (Tremadocian)

 Argentina

A member of Olenidae.

Parabolinella pompadouris[146]

Sp. nov

Valid

Monti, del Huerto Benítez & Ramirez

Ordovician (Tremadocian)

 Argentina

A member of Olenidae.

Paraperaspis striolatus[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Paraphillipsinella ciliensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Pararemopleurides[127]

Gen. et 3 sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Genus includes new species P. longispinus, P. schizophyllus and P. shaanxiensis.

Parashanxiella[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Proasaphiscidae.

Paraszeaspis[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Ignotogregatidae.

Parillaenus xui[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Pelitlina? corbachoi[125]

Sp. nov

Valid

Basse & Müller

Early Devonian

 Spain

Perizostra[133]

Gen. et sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

A member of Scutelluidae. Genus includes new species P. campbelli.

Phacops boudjemaai[132]

Sp. nov

Valid

Khaldi et al.

Early Devonian

‘Chefar el Ahmar’ Formation

 Algeria

A phacopid. Originally described as a species of Phacops; Van Viersen, Holland & Koppka (2017) transferred the species to the genus Adrisiops.[135]

Platypeltoides hammondi[147]

Sp. nov

Valid

Corbacho & López-Soriano

Ordovician (late Tremadocian)

Fezouata formation

 Morocco

A nileid.

Poulsenia venusta[126]

Sp. nov

Valid

Pegel in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Ptychopariidae.

Prolisania[130]

Gen. nov

Valid

Yuan & Zhu in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Lisaniidae.

Pseudopetigurus yuandunensis[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Psychopyge angeles[125]

Sp. nov

Valid

Basse & Müller

Devonian (Emsian)

 Morocco

Qianlishania[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Solenopleuridae.

Quintonia[133]

Gen. et 2 sp. nov

Valid

Holloway & Lane

Silurian

Tomcat Creek limestone

 Australia

A member of Scutelluidae. Genus includes new species Q. arata and Q. pavo.

Rasettiella[148]

Gen. et comb. et 2 sp. nov

Junior homonym

Wernette & Westrop

Cambrian

 Canada

A dolichometopid. Genus includes species previously assigned to the genus Bathyuriscidella, as well as Rasettiella lupiae and Rasettiella sorgehanarum. The generic name is preoccupied by Rasettiella Özdikmen (2005).

Rhenocynproetus vanvierseni[125]

Sp. nov

Valid

Basse & Müller

Devonian (Emsian)

Rupbach Shale

 Germany

Rickardsia[149]

Gen. et comb. nov

Valid

Storey, Thomas & Owens

Silurian

 Czech Republic
 Sweden
 United Kingdom

A member of Dalmanitidae. The type species is "Dalmanites" mobergi Hede (1915); genus also includes "Delops" dermolac Šnajdr (1981) and "Phacops (Dalmania)" nobilis Thomas (1900).

Rina? mayskaya[136]

Sp. nov

Valid

Bushuev & Makarova

Cambrian

Chaya Formation

 Russia

A member of Solenopleuridae.

Rorringtonia conica[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Rorringtonia minuta[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Rorringtonia obsoleta[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Rotundampyx[123]

Gen. et sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

A raphiophorid. Genus includes new species R. lasengmiaoensis.

Ryaneyreus[129]

Gen. et sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

A member of the family Parabolinoididae. Genus includes news species R. zoeae.

Sagavia obscura[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Scabriscutellum (Rheiscutellum)[125]

Subgen. nov

Valid

Basse & Müller

Devonian (Emsian)

Rupbach Shale

 Germany

Sciaspis[130]

Gen. nov

Valid

Zhu in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Ordosiidae.

Sibirolenus[124]

Gen. et comb. nov

Valid

Geyer

Cambrian

 Russia

A member of Palaeolenidae; a new genus for "Micmacca" enormis.

Sinoanomocare[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Anomocaridae.

Strictaspis[126]

Gen. et sp. nov

Valid

Lazarenko & Salikhova in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Asaphiscidae. The type species is S. leve.

Sulcipagetia[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Eodiscidae.

Taenicephalus leei[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Taenicephalus westropi[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Tanglefootia[129]

Gen. et sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

A member of the family Cheilocephalidae. Genus includes new species T. calvini.

Taosigouia[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Anomocarellidae.

Telephina truncata[127]

Sp. nov

Valid

Zhou, Zhou & Xiang

Ordovician

Pagoda Formation

 China

Telopeltis intermedia[145]

Sp. nov

Valid

McNamara & Feist

Devonian (Frasnian)

Virgin Hills Formation

 Australia

A species of Telopeltis.

Tomagnostus apertus[143]

Sp. nov

Valid

Makarova & Bushuev

Cambrian

Chaya Formation

 Russia

A member of Ptychagnostidae.

Tongxinaspis ovata[123]

Sp. nov

Valid

Zhou, Zhou & Yin

Ordovician (Darriwilian)

Zhuozishan Formation

 China

A dionidid.

Torleyiscutellum[150]

Gen. et sp. nov

Valid

Basse, Koch & Lemke

Devonian (early Givetian)

 Germany

A scutelluid trilobite. Genus includes new species T. herwigorum.

Trifoliops cronierae[137]

Sp. nov

Valid

Feist in Feist, Mahboubi & Girard

Late Devonian

“Argiles de Marhouma” Formation

 Algeria

A member of Phacopidae.

Trimerocephalus mahboubii[137]

Sp. nov

Valid

Feist in Feist, Mahboubi & Girard

Late Devonian

“Argiles de Marhouma” Formation

 Algeria

A member of Phacopidae.

Tropidocoryphe werneri[125]

Sp. nov

Valid

Basse & Müller

Devonian (Emsian)

Rupbach Shale

 Germany

Ulakhanella triangula[126]

Sp. nov

Valid

Pegel in Pegel et al.

Cambrian

 Russia

A member of Ptychopariida belonging to the family Palaeolenidae.

Verkholenoides cognatus[126]

Sp. nov

Valid

Pegel & Salikhova in Pegel et al.

Cambrian

 Russia

A member of Asaphida belonging to the family Pterocephaliidae.

Vicnepea[144]

Gen. nov

Valid

Bentley, Jago & Cooper

Cambrian (Drumian)

Antarctica
(Victoria Land)

A polymerid trilobite.

Wuhaiaspis[130]

Gen. nov

Valid

Yuan & Zhang in Yuan, Zhang & Zhu

Cambrian

 China

A member of the family Ptychopariidae.

Wujiajiania lyndasmithae[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Wujiajiania ricksmithi[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

Other arthropods

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Acanthomeridion anacanthus[156]

Sp. nov

Valid

Hou et al.

Early Cambrian

 China

Agnostotes weugi[129]

Sp. nov

Valid

Chatterton & Gibb

Cambrian (Furongian)

 Canada

A member of Agnostida.

Aquilonifer[157][158][159]

Gen. et sp. nov

Valid

Briggs et al.

Silurian (late Wenlock)

Herefordshire Lagerstätte

 United Kingdom

A stem-mandibulate. The type species is Aquilonifer spinosus.

Archeallacma[160]

Gen. et sp. nov

Valid

Sánchez-García & Engel[161]

Early Cretaceous

 Spain

A springtail belonging to the family Sminthuridae. The type species is A. dolichopoda.

Brachyaglaspis[162]

Gen. et sp. nov

Valid

Ortega-Hernández, Van Roy & Lerosey-Aubril

Ordovician (middle Floian)

Fezouata Konservat-Lagerstätte

 Morocco

A member of Aglaspidida. The type species is Brachyaglaspis singularis.

Burmisotoma spinulifera[163]

Sp. nov

Valid

Sánchez-García & Engel

Early Cretaceous (late Albian)

Utrillas Group

 Spain

A springtail belonging to the family Isotomidae.

Clypecaris serrata[164]

Sp. nov

Valid

Yang et al.

Cambrian

Hongjingshao Formation

 China

A member of Deuteropoda of uncertain phylogenetic placement.

Cretokatianna[160]

Gen. et sp. nov

Valid

Sánchez-García & Engel[161]

Early Cretaceous

 Spain

A springtail belonging to the family Katiannidae. The type species is C. bucculenta.

Decoracaris[50]

Gen. et sp. nov

Valid

Briggs et al.

Ordovician (Darriwilian)

Winneshiek Lagerstätte

 United States

A bivalved arthropod of uncertain phylogenetic placement, possibly a thylacocephalan. The type species is Decoracaris hildebrandi.

Dyrnwynia[165]

Gen. et sp. nov

Valid

Legg

Ordovician (middle Darriwilian)

Llanfallteg Formation

 United Kingdom

A member of Marrellida. The type species is Dyrnwynia conollyi.

Enosiaspis[166]

Gen. et sp. nov

Valid

Legg

Ordovician (Tremadocian)

Fezouata Formation

 Morocco

A marrellomorph belonging to the group Acercostraca. The type species is Enosiaspis hrungnir.

Hipponicharion pearylandica[167]

Sp. nov

Valid

Peel et al.

Cambrian

Henson Gletscher Formation

 Greenland

A member of Bradoriida. Originally described as a species of Hipponicharion; subsequently made the type species of a separate genus Navarana.[168]

Iosuperstes[50]

Gen. et sp. nov

Valid

Briggs et al.

Ordovician (Darriwilian)

Winneshiek Lagerstätte

 United States

A bivalved arthropod of uncertain phylogenetic placement. The type species is Iosuperstes collisionis.

Katiannasminthurus[160]

Gen. et sp. nov

Valid

Sánchez-García & Engel[161]

Early Cretaceous

 Spain

A springtail, possibly a member of the family Sminthuridae. The type species is K. xenopygus.

Liangshanella? nivalis[167]

Sp. nov

Valid

Peel et al.

Cambrian

Henson Gletscher Formation

 Greenland

A member of Bradoriida.

Paleolimulus woodae[169]

Sp. nov

Valid

Lerner, Lucas & Mansky

Carboniferous (Tournaisian)

Horton Bluff Formation

 Canada

A member of Xiphosura belonging to the family Paleolimulidae. Originally described as a species of Paleolimulus, but subsequently transferred to the genus Norilimulus.[170]

Proisotoma communis[163]

Sp. nov

Valid

Sánchez-García & Engel

Early Cretaceous (late Albian)

Utrillas Group

 Spain

A springtail belonging to the family Isotomidae.

Protoisotoma autrigoniensis[163]

Sp. nov

Valid

Sánchez-García & Engel

Early Cretaceous (late Albian)

Utrillas Group

 Spain

A springtail belonging to the family Isotomidae.

Pseudosminthurides[160]

Gen. et sp. nov

Valid

Sánchez-García & Engel[161]

Early Cretaceous

 Spain

A springtail belonging to the family Sminthurididae. The type species is P. stoechus.

Skryjagnostus usitatus[126]

Sp. nov

Valid

Salikhova in Pegel et al.

Cambrian

 Russia

A member of Agnostida (a group of arthropods of uncertain phylogenetic placement, possibly trilobites).

Sphyrotheciscus[160]

Gen. et sp. nov

Valid

Sánchez-García & Engel[161]

Early Cretaceous

 Spain

A springtail belonging to the family Sminthuridae. The type species is S. senectus.

Tomagnostus brantevikensis[171]

Sp. nov

Valid

Weidner & Nielsen

Middle Cambrian

Alum Shale Formation

 Sweden

A member of Agnostida belonging to the family Ptychagnostidae.

Wisangocaris[172]

Gen. et sp. nov

Valid

Jago, García-Bellido & Gehling

Early Cambrian

Emu Bay Shale

 Australia

A member of Chelicerata sharing many overall body features with Sanctacaris. The type species is Wisangocaris barbarahardyae.

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