Domestic yak

The domestic yak (Bos grunniens) is a long-haired domesticated cattle found throughout the Himalayan region of the Indian subcontinent, the Tibetan Plateau, Northern Myanmar, Yunnan, Sichuan and as far north as Mongolia and Siberia. It is descended from the wild yak (Bos mutus).[1]

"Yak" redirects here. For the related species Bos mutus, see Wild yak. For other uses, see Yak (disambiguation).

Yak
A yak in the Nepalese Himalayas.
Domesticated
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Artiodactyla
Family: Bovidae
Subfamily: Bovinae
Genus: Bos
Species:
B. grunniens
Binomial name
Bos grunniens
Linnaeus, 1766

Etymology

The English word "yak" is a loan originating from Tibetan: གཡག་, Wylie: g.yag . In Tibetan and Balti it refers only to the male of the species, the female being called Tibetan: འབྲི་, Wylie: 'bri or Tibetan: གནག, Wylie: g.nag in Tibetan and Tibetan: ཧཡག་མོ་, Wylie: hYag-mo in Balti. In English, as in most other languages that have borrowed the word, "yak" is usually used for both sexes, with "bull" or "cow" referring to each sex separately.

Taxonomy
Bronze yak, Yuan dynasty

Yaks belong to the genus Bos and are therefore related to cattle (Bos primigenius species). Mitochondrial DNA analyses to determine the evolutionary history of yaks have been inconclusive.

The yak may have diverged from cattle at any point between one and five million years ago, and there is some suggestion that it may be more closely related to bison than to the other members of its designated genus.[2] Apparent close fossil relatives of the yak, such as Bos baikalensis, have been found in eastern Russia, suggesting a possible route by which yak-like ancestors of the modern American bison could have entered the Americas.[3]

The species was originally designated as Bos grunniens ("grunting ox") by Linnaeus in 1766, but this name is now generally considered to refer only to the domesticated form of the animal, with Bos mutus ("mute ox") being the preferred name for the wild species. Although some authors still consider the wild yak to be a subspecies, Bos grunniens mutus, the ICZN made an official ruling in 2003[4] permitting the use of the name Bos mutus for wild yaks, and this is now the more common usage.[5][3][6]

Except where the wild yak is considered as a subspecies of Bos grunniens, there are no recognised subspecies of yak.

Physical characteristics
A domestic yak at Yamdrok Lake.

Yaks are heavily built animals with bulky frames, sturdy legs, rounded, cloven hooves, and extremely dense, long fur that hangs down lower than the belly. While wild yaks are generally dark, blackish to brown in colouration, domestic yaks can be quite variable in colour, often having patches of rusty brown and cream. They have small ears and wide foreheads, with smooth horns that are generally dark in colour. In males (bulls), the horns sweep out from the sides of the head, and then curve forward. They typically range from 48 to 99 cm (19 to 39 in) in length. The horns of females (cows) are smaller, only 27 to 64 cm (11 to 25 in) in length, and have a more upright shape. Both sexes have a short neck with a pronounced hump over the shoulders, although this is larger and more visible in males.[3] Males weigh 350 to 585 kg (772 to 1,290 lb), females weigh 225 to 255 kg (496 to 562 lb). Wild yaks can be substantially heavier, bulls reaching weights of up to 1,000 kilograms (2,200 lb).[7] Depending on the breed, domestic yak males are 111–138 centimetres (44–54 in) high at the withers, while females are 105–117 centimetres (41–46 in) high at the withers.[8]

Both sexes have long shaggy hair with a dense woolly undercoat over the chest, flanks, and thighs to insulate them from the cold. Especially in bulls, this may form a long "skirt" that can reach the ground. The tail is long and horselike rather than tufted like the tails of cattle or bison. Domesticated yaks have a wide range of coat colours, with some individuals being white, grey, brown, roan or piebald. The udder in females and the scrotum in males are small and hairy, as protection against the cold. Females have four teats.[3]

Yaks are not known to produce the characteristic lowing (mooing) sound of cattle, but both wild and domestic yaks grunt and squeak, which inspired the scientific name of the domestic yak variant, Bos grunniens (grunting bull). Nikolay Przhevalsky named the wild variant Bos mutus (silent bull) believing that it did not make a sound at all, but it does.[9]

Physiology
Yak rider near Tsomgo Lake, Sikkim (3700 m)

Yak physiology is well adapted to high altitudes, having larger lungs and heart than cattle found at lower altitudes, as well as greater capacity for transporting oxygen through their blood[10] due to the persistence of foetal haemoglobin throughout life.[11] Conversely, yaks have trouble thriving at lower altitudes,[12] and are prone to suffering from heat exhaustion above about 15 °C (59 °F). Further adaptations to the cold include a thick layer of subcutaneous fat, and an almost complete lack of functional sweat glands.[10]

Compared with domestic cattle, the rumen of yaks is unusually large, relative to the omasum. This likely allows them to consume greater quantities of low-quality food at a time, and to ferment it longer so as to extract more nutrients.[10] Yak consume the equivalent of 1% of their body weight daily while cattle require 3% to maintain condition.

Odour

Contrary to popular belief, yak and their manure have little to no detectable odour[13] when maintained appropriately in pastures or paddocks with adequate access to forage and water. Yak's wool is naturally odour resistant.[14]

Reproduction and life history
Ten-day-old yak.

Yaks mate in the summer, typically between July and September, depending on the local environment. For the remainder of the year, many bulls wander in small bachelor groups away from the large herds, but, as the rut approaches, they become aggressive and regularly fight among each other to establish dominance. In addition to non-violent threat displays, bellowing, and scraping the ground with their horns, bull yaks also compete more directly, repeatedly charging at each other with heads lowered or sparring with their horns. Like bison, but unlike cattle, males wallow in dry soil during the rut, often while scent-marking with urine or dung.[3] Females enter oestrus up to four times a year, and females are receptive only for a few hours in each cycle.[15]

Gestation lasts between 257 and 270 days,[10] so that the young are born between May and June, and results in the birth of a single calf. The cow finds a secluded spot to give birth, but the calf is able to walk within about ten minutes of birth, and the pair soon rejoin the herd.[10] Females of both the wild and domestic forms typically give birth only once every other year,[3] although more frequent births are possible if the food supply is good.

Calves are weaned at one year and become independent shortly thereafter. Wild calves are initially brown in color, and only later develop the darker adult hair. Females generally give birth for the first time at three or four years of age,[16] and reach their peak reproductive fitness at around six years. Yaks may live for more than twenty years in domestication or captivity,[3] although it is likely that this may be somewhat shorter in the wild.

Hybrid yak

In Nepal, Tibet and Mongolia, domestic cattle are crossbred with yaks. This gives rise to the infertile male dzo མཛོ། as well as fertile females known as མཛོ་མོ། dzomo or zhom, which may be crossed again with cattle. The "Dwarf Lulu" breed, "the only Bos primigenius taurus type of cattle in Nepal" has been tested for DNA markers and found to be a mixture of both taurine and zebu types of cattle (B. p. taurus and B. p. indicus) with yak.[17] According to the International Veterinary Information Service, the low productivity of second generation cattle-yak crosses makes them suitable only as meat animals.[18]

Crosses between yaks and domestic cattle (Bos primigenius taurus) have been recorded in Chinese literature for at least 2,000 years.[3] Successful crosses have also been recorded between yak and American bison,[18] gaur, and banteng, generally with similar results to those produced with domestic cattle.[3]

Relationship with humans

Domesticated yaks have been kept for thousands of years, primarily for their milk, fibre and meat, and as beasts of burden. Their dried droppings are an important fuel, used all over Tibet, and are often the only fuel available on the high treeless Tibetan Plateau. Yaks transport goods across mountain passes for local farmers and traders as well as for climbing and trekking expeditions. "Only one thing makes it hard to use yaks for long journeys in barren regions. They will not eat grain, which could be carried on the journey. They will starve unless they can be brought to a place where there is grass."[19] They also are used to draw ploughs.[20] Yak's milk is often processed to a cheese called chhurpi in Tibetan and Nepali languages, and byaslag in Mongolia. Butter made from yaks' milk is an ingredient of the butter tea that Tibetans consume in large quantities,[21] and is also used in lamps and made into butter sculptures used in religious festivities.[22]

Yak racing

Husbandry research

The Indian government established a dedicated centre for research into yak husbandry, the ICAR-National Research Centre on Yak, in 1989. It is located at Dirang, Arunachal Pradesh, and maintains a yak farm in the Nyukmadung area at an altitude of 2,750 metres (9,020 ft) above MSL.[23]

Yak sports

In parts of Tibet and Karakorum, yak racing is a form of entertainment at traditional festivals and is considered an important part of their culture. More recently, sports involving domesticated yaks, such as yak skiing or yak polo, are being marketed as tourist attractions in Central Asian countries, including in Gilgit-Baltistan, Pakistan.

See also

References
  1. Grubb, P. (2005). "Order Artiodactyla". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 691. ISBN 978-0-8018-8221-0. OCLC 62265494.
  2. Guo, S.; et al. (2006). "Taxonomic placement and origin of yaks: implications from analyses of mtDNA D-loop fragment sequences". Acta Theriologica Sinica. 26 (4): 325–330.
  3. Leslie, D.M.; Schaller, G.B. (2009). "Bos grunniens and Bos mutus (Artiodactyla: Bovidae)". Mammalian Species. 836: 1–17. doi:10.1644/836.1.
  4. International Commission on Zoological Nomenclature (2003). "Opinion 2027. Usage of 17 specific names based on wild species which are predated by or contemporary with those based on domestic animals (Lepidoptera, Osteichthyes, Mammalia): conserved". Bulletin of Zoological Nomenclature. 60: 81–84.
  5. Harris, R.B.; Leslie, D. (2008). "Bos mutus". IUCN Red List of Threatened Species. 2008. Retrieved 4 September 2014.
  6. Gentry, A.; Clutton-Brock, J.; Groves, C. P. (2004). "The naming of wild animal species and their domestic derivatives". Journal of Archaeological Science. 31 (5): 645. doi:10.1016/j.jas.2003.10.006.
  7. Buchholtz, C. (1990). True Cattle (Genus Bos). pp. 386–397 in S. Parker, ed. Grzimek's Encyclopedia of Mammals, Volume 5. New York: McGraw-Hill Publishing Company. (quoted in Oliphant, M. (2003). Bos grunniens (On-line), Animal Diversity Web. Accessed 4 April 2009)
  8. "The Yak. Chapter 2: Yak breeds". FAO. Retrieved 31 August 2017.
  9. "Origins, Domestication and Distribution of Yak". fao.org. Retrieved 31 August 2017.
  10. Wiener, Gerald; Jianlin, Han; Ruijun, Long (2003). "4 The Yak in Relation to Its Environment", The Yak, Second Edition. Bangkok: Regional Office for Asia and the Pacific Food and Agriculture Organization of the United Nations, ISBN 92-5-104965-3. Accessed 8 August 2008.
  11. Sarkar, M.; Das, D. N.; Mondal, D. B. (1999). "Fetal Haemoglobin in Pregnant Yaks (Poephagus grunniens L.)". The Veterinary Journal. 158 (1): 68–70. doi:10.1053/tvjl.1999.0361. PMID 10409419.
  12. Yak, Animal genetics training resources version II: Breed Information. Adopted from: Bonnemaire, J. "Yak". In: Mason, Ian L. (ed). (1984). Evolution of Domesticated Animals. London: Longman, pp. 39–45. ISBN 0-582-46046-8. Accessed 8 August 2008.
  13. Yak Dung. Sherpatrek.com. Retrieved on 2012-12-19.
  14. "Superior Properties of Yak Wool". Archived from the original on 6 December 2010. Retrieved 3 May 2012.CS1 maint: bot: original URL status unknown (link)
  15. Sarkar, M.; Prakash, B.S. (2005). "Timing of ovulation in relation to onset of estrus and LH peak in yak (Poephagus grunniens L.)". Animal Reproduction Science. 86 (4): 353–362. doi:10.1016/j.anireprosci.2004.08.005. PMID 15766812.
  16. Zi, X.D. (2003). "Reproduction in female yaks (Bos grunniens) and opportunities for improvement". Theriogenology. 59 (5–6): 1303–1312. doi:10.1016/S0093-691X(02)01172-X. PMID 12527077.
  17. Takeda, K.; Satoh, M.; Neopane, S.P.; Kuwar, B.S.; Joshi, H.D.; Shrestha, N.P.; Fujise, H.; Tasai, M.; Tagami, T.; Hanada, H. (2004). "Mitochondrial DNA analysis of Nepalese domestic dwarf cattle Lulu". Animal Science Journal. 75 (2): 103. doi:10.1111/j.1740-0929.2004.00163.x.
  18. Zhang, R.C. (14 December 2000). "Interspecies Hybridization between Yak, Bos taurus and Bos indicus and Reproduction of the Hybrids". In: Recent Advances in Yak Reproduction, Zhao, X.X.; Zhang, R.C. (eds.). International Veterinary Information Service.
  19. Golden Book Encyclopedia, Vol. 16 p. 1505b. Rockefeller Center, NY: Golden Press (1959).
  20. Gyamtsho, Pema. "Economy of Yak Herders" (PDF).
  21. Tibet and Tibetan Foods. Flavorandfortune.com. Retrieved on 2012-12-19.
  22. Yaks, butter & lamps in Tibet, webexhibits.org
  23. "ICAR-National Research Centre on Yak".
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